1907.] OF A FROG OF THE GENUS MEGALOPHRYS. 331 



on each shoulder and one on sacral region. Verbebrfe proccelous ; 

 sacral vertebra fused with coccyx, its transverse processes greatly 

 expanded. No ribs. Omosternum not rudimentary, with calcified 

 style; sternum a bony style, with expanded cai'tilaginous xiphi- 

 sternum ; shoulder-girdle arciferous. Fingers free ; inner finger 

 longer than second. Toes half-webbed ; outer toe separated from 

 next by web. Toes not dilated at tips ; no articular tubercles ; 

 inner metatarsal tubercle extensive but not separated anteriorly 

 from the surface of the foot ; no outer metatarsal tubercle. 



Hah. Borneo. 



It will be observed that in the above brief diagnosis I have not 

 attempted to differentiate between generic aiid specific characters 

 or indeed family characters. That this Frog belongs to the 

 Pelobatidae is quite plain. That it cannot be referred to any 

 known genus — if the present definitions of the same are retained — 

 is also obvious*. I propose, however, for the present to defer the 

 question of generic distinctness until the Pelobatidte are better 

 known ana,tomically f. 



Whether this Frog is a new species or identical with Megalophrys 

 nasuta I must, for reasons already stated, leave uncertain. 



§ Mitscles of the Back. 



In the various drawings (text-figs. 93-96) submitted herewith in 

 illustration of the diaphragm of different species of Batrachia, the 

 principal muscles of the back are also shown, and it will be at 

 once seen that these difier considerably in the different forms. 

 Only in Megalophrys is it impossible to see these different muscles 

 (text-fig. 98), for the lengthy transverse muscle of the oesophagus 

 and lung almost completely covers them over. When, however, 

 this muscle is divided along its greater length, i. e. across the 

 direction of its fibres and the two flaps reflected, the muscles now in 

 question are fully displayed as shown in text -fig. 93. The same 

 muscles are present which are known to exist in Rana escidenta, 

 with the possible addition of a muscle not found either in 

 R. escidenta, R. guppyi (see p. 333), or R. tigrina, of which species 

 I have examined the last two. The Intertra^isversarii muscles 

 are well developed and commence from the anterior margin 

 of the greatly expanded transverse process of the sacral vertebrfe. 

 They are so well developed that they almost conceal the anteriorly 

 lying transverse processes of the vertebrae, each separate band of 

 muscle joining successive transverse processes being attached near 

 to the ventral median line of the same. The Ilio-lumbaris muscle 



* (Footnote added May 8th.) After the reading of tliis paper, Mr. Boulenger 

 kindly drew ray attention to the fact that there occurs m Felohates cultripes 

 a variation in connexion with the fusion or nou-fusion of the sacral vertebra 

 with the coccyx ('The Tailless Batrachia of Europe,' Raj' Soc. 1897, part i. p. 208). 

 It does not, however, follow that if a character is variable in one species it is not 

 a " good " character in another. 



t Schlegel's name Ceratophryne cannot be resuscitated ; for he first applied it to 

 C. dorsata, au American frog which is, I presume, Ceratoplirys dorsata. 



