688 MR. F. E. BEDDARD ON THE [June 15, 



gizzard-like pylorus was at least 12 mm., a thickness which is 

 quite as great as that of the other Anteater, I imagine. 



The hejxtto-gastric ligament or lesser omentiim presents one or 

 two features of interest. The greater part of this ligament was, 

 as in other mammals, horizontal in direction, roofing-over that 

 section of the coelom which communicates with the larger section 

 of the abdominal cceloni by the foramen of Winslow. The edge 

 of this hepato-gastric ligament, lying just over the foramen of 

 Winslow, was raised into a deep vertical membrane fully three- 

 quai-ters of an inch deep, which passes dorsally of the right central 

 lobe of the liver. This is of course all part of the lesser omentum 

 or hepato-gastric ligament ; but the vertical part seems to me to 

 represent the primitive ventral mesentery, connected directly with 

 the middle line of the ventral parietes and not via the liver. The 

 greater omentum is but little developed and, as Flower has 

 mentioned, is not attached to the colon anywhere ; it is indeed 

 not visible when the animal is opened for dissection, being 

 covered by rather than covering the coils of the intestines. The 

 splenic omentum is short, and just laps loosely over the pancreas. 

 It can be stretched out straight quite easily, and is inserted 

 on to the oesophageal border of the stomach, along which runs 

 one of the important gastric branches of the portal. It 

 connects the pancreas and the spleen with this region of the 

 stomach. A remarkable fact about the stomach of this Edentate 

 is the large number of lymphatic glands which are found in the 

 region of the stomach and, as I shall explain later, of the 

 intestine. As to those which occur in the region of the stomach, 

 there are first of all three, one above the other, which lie on the- 

 side of the junction between duodenum and stomach — that side 

 which faces the abdominal cavity. These glands are in close con- 

 nection with the portal branch, which passes from the oesophageal 

 border of the stomach to join the main portal trunk as it passes 

 dorsally to the duodenum on its way to the liver. Two other 

 lymphatic glands lie in the lesser omentum, one near to the 

 oesophagus and the other close to the vertical membrane already 

 described. Finally there are two other lymj^hatic glands on the 

 pancreas side of the vertical membrane — to the right, therefore, 

 of the expanse of membrane which connects the stomach with the 

 liver and may be termed, and is termed, lesser omentum. An 

 eighth gland is partly imbedded in the tip of the pancreas, where 

 it runs towards the liver in close connection with the cystic duct 

 of the gall-bladder. 



The Small Intestine in my specimen measured at least 97 inches 

 in length. This is rather understating the length, I imagine, 

 for I was anxious not to stretch it unduly. Sir W. Flower gives 

 100 inches as the length of the small intestine in the example 

 measured by himself. In the latter example of Tamandua the 

 colon was only 5 ^ inches long ; I found that from the caecum to 

 the anus was quite 7 inches in my specimen. Sir W. Flower 

 remarks that the duodenum has no mesentery attaching it to the 



