842 Transactions of the Society. 



absence of an envelope to the fructification, and in the carpogonium 

 remaining undivided. 



I have much pleasure in naming the genus after my friend 

 Mr. J. T. Milow, to whom I am indebted for many rare and 

 interesting fungi. 



The sporidia of this fungus were sown on a glass slip, moistened 

 with a mixture of glycerin and the liquid from decaying rushes, 

 placed under a bell-jar, and kept in the dark. After a few days 

 some of the sporidia showed a slight projection at one end, the 

 exospore split at this point, and the endospore protruded as a 

 hypha. Out of some hundreds of sporidia that germinated only 

 four were observed to give origin to more than one thread, these 

 produced two, one springing from each end. Immediately pre- 

 ceding germination the protoplasm becomes granular and opaque, 

 and contains several large refractive globules of an oily-looking 

 substance ; after germination the whole of the contents pass into 

 the mycelium, which is cut off from the cavity of the sporidium 

 by a transverse septum formed close to the latter. The hyphae 

 after elongating for some distance as simple threads with but few 

 septa, branch repeatedly in a monopodial manner, these in turn 

 give off numerous lateral branches in acropetal succession ; some 

 resemble the branch from which they spring, in having cells 

 about four times as long as broad, and are probably organs of 

 nutrition, penetrating the tissues of the host, and undergoing no 

 further modification of form ; others, which eventually give origin 

 to the organs of reproduction, may be recognized during the 

 earliest stages of development, by the much shorter cells, not 

 longer than broad, and invariably three in number ; of these the 

 terminal one becomes the carpogonium, the basal one gives origin 

 to the pollinodium as a lateral branch, while the central one re- 

 mains unchanged, and forms the basal cell of the fruit. In some 

 instances the poUinodium is not developed ; then the terminal cell 

 of the fertile branch, instead of forming a carpogonium, elongates 

 and gives origin to a chain of conidia ; this asexual form of repro- 

 duction occurs mixed with the sexual form. 



The three cells forming the fertile branch are at first of equal 

 size, but the terminal one, owing to continued growth, soon becomes 

 very much larger than the remaining two, assumes a broadly 

 obovate form, and contains coarsely granular opaque protoplasm, 

 crowded with vacuoles of varying size ; this cell is the carpogonium, 

 now ready for fertilization. During the development of the car- 

 pogonium, the pollinodium originates as a lateral outgrowth from 

 the basal cell of the reproductive branch ; this outgrowth, during 

 elongation, assumes a clavate form, at the same time curving 

 upwards towards the carpogonium; a well-defined nucleus con- 



