Internal Anatomy of Urojpoda Krameri. By A.D. Michael. 9 



The annular form of the system, taken as a whole, which is so well 

 known in the Arachnida, and which is so conspicuous in that of the 

 Orihatidse, is equally clearly shown in the female of Uropoda 

 Krameri; but in the male this form is more lost, in consequence 

 of the absence of the long vasa deferentia which form an element of 

 the ring in the Orihatidm. Probably it is only those who know how 

 the ring is formed in the males of the last-named family who 

 would recognize some vestige of it in those of the Uropoda. 



The Male. Figs. 10-13. 



The male organs lie almost immediately below the ventriculus ; 

 they consist of a central chamber, six more or less sack-like organs, 

 and a large single duct leading to the penis. The most conspicuous 

 of these is the central chamber (vs), a large heart-shaped organ 

 compressed dorso-ventrally, and having the broader end turned 

 forward; this organ is the nearest to the ventral level, the other 

 parts of the system lying slightly above it. I take it to be partly 

 glandular in its office, and also to some extent to function as a vesicula 

 seminalis ; in which case it would agree with the corresponding organ 

 in the Orihatidm ; and this appears to be Winkler's view with 

 regard to the organ in his species, TJ. ohsoura ; in which case, how- 

 ever, the organ appears from his description to be more globular. 



Four long, sack-like, glandular organs (t^, t^) take their origin 

 immediately above the central chamber, and near its anterior margin. 

 They do not appear to communicate directly with the central chamber, 

 but all seem to open into a small median antechamber. The sacks 

 are pyriform, smallest where they enter the antechamber, and largest 

 at the blind, free ends. One pair, which are usually somewhat the 

 larger, are nearly straight, and are directed almost backward. The 

 corresponding organs in U. ohscura are regarded by Winkler as being 

 the true testes. The second pair, the mouths of which are placed 

 above those of the first pair, are more curved, or comma-shaped; 

 they are directed almost transversely across, and partly under the 

 central chamber ; their distal ends curve backward. If these cor- 

 respond to the second pair of sacks figm-ed by Herr Winkler in his 

 diagram he regards them as accessory glands, not testes ; but as he 

 only mentions four sacks in his species, and I find six in mine, it is 

 probable that his accessory glands correspond to the smaller sacks 

 (oil-glands) mentioned immediately below, and that the pair of organs 

 now treated of are rather to be regarded as a second pair of testes ; at 

 all events they greatly resemble the first pair. In addition to these 

 four there are the two other sacks above referred to (g), they are 

 much smaller and almost globular. These organs have thin walls, 

 and contain only a highly refractive oily liquid. They are placed one 

 on each side of the ductus ejaculatorius, and apparently communicate 

 with the small median antechamber. Somewhat similar organs exist 

 in a few of the Orihatidw, but not in all. 



