AXATOMY OF THE TAKIN. 217 



limb of the saprasjlvian sulcus. These last two features are not 

 constant iu the ovine brain, however, and are not seen in the 

 sheep brain figured in the Catalogue of the Museum of the 

 Royal College of Surgeons. 



Where the brain of the Takin difiers from that of the sheep 

 it agrees with the bovine type represented by the Anoa. The 

 arrangement of orbital, anterior terminal and rhinal, and the 

 relative positions of the transverse, coronal and suprasylvian sulci 

 are almost identical in these two, but the cruciate sulcus is even 

 less marked in the Anoa than in the sheep. The development of 

 this sulcus seems to be partly associated with the position of the 

 coronal sulcus with i-eference to the supero-mesial margin. In 

 the Anoa the gyrus between them is less than 3 mm. in width, 

 in the sheep 7 mm., but in the Takin 16 mm. There is thus room 

 for a long "cruciate" sulcus on the lateral aspect in the Takin, 

 for a very short one only in the sheep, and in the bovine brain 

 the cruciate upturning is confined to the mesial aspect. 



Baclorcas differs markedly from the bovine type in the absence 

 of any approach to the curious " double" form of Sylvian fissure 

 found in the ox-like brain. 



. The anterior corpus quadrigeminum is as much larger in the 

 Takin than in the sheep as it is larger in the sheep than in the 

 Anoa, but the difference is slight and may be partly due to 

 fiattening consequent upon preservation. The body is distinctly 

 triangular in outline in mesial section in the Takin, oval in the 

 sheep, and more flattened still in the Anoa. 



A similar progressive change is seen in the relative position of 

 the supra-callosal gyrus and splenium. In the Anoa there is 

 hardly any cortex directly behind the splenium ; the whole mass 

 lies underneath the corpus callosum ; in Ovis the small pointed 

 extremity of the hippocampus nudus lies in a line with the middle 

 of the splenium,- iu Baclorcas, -^fi stated, the gyrus is 5 mm. broad 

 horizontally behind the splenium. 



The occipital pole is more pointed and projects rather further 

 backwards over the cerebellum in the Takin and Anoa than in 

 the sheep. 



It seems hardly advisable to attempt to draw far-reaching 

 conclusions of systematic importance from the scanty material at 

 present available, more especially since there are no data on record 

 of the anatomy of the other members of the Rupricaprine section. 



Support is given to those wdio hold the aflinity of Biodorcas 

 with Ovibos to be a close one ; the two animals d.iffer in but few 

 points of their soft anatomy, and in many cases they share 

 characters which differentiate them from other rummaiit 

 Artiodactyla . 



The suggested relationship between Connochcetes and Budorcas 

 is not borne out ; the resemblances between the two animals are 

 few and far between and are mostly points of detail and minor 

 importance. 



Between Bos and Ovis, Budorcas seems to hold an intermediate 

 Paoc. ZooL. See— 1919, No. XV. 15 



