ON EXTERNAL CHARACTERS OF THE PARADOXURIN^. 387 



30. On the Feet and Grlands and other External Characters 

 of the Paradoxurine Genera Paradoxurus, Arctictis, 

 Arctogalidia, and Nandinia. By H. I. PocoCK, F.R.S., 

 F.L.S., F.Z.S., Curator o£ Mammals. 



[Received May 11, 1915 : Read June 8, 1915.] 

 (Text-figures 1-10.) 



I^i'^^- Page 



Feet of Paradoxurus 387 



„ Arctictis 389 



„ Arctoffalidia 390 



„ Nandinia 392 



Comparison between the Paradoxurine and Viverrine types 



of feet 394 



Rhinariuni and vibrissa3 of Paradoxurines 395 



Ear of Paradoxurines and Viverrines 397 



Perfume-gland and external genitalia of Faradoxurus ... 401 



,, „ „ „ Arctictis 406 



„ „ „ „ Arctogalidia ... 407 



„ „ ., „ Nandinia 409 



General conclusions with regard to the perfume-glaiids ... 411 



This paper, the result of researches carried on at intervals for 

 several years in the Society's Prosectorial Department, may be 

 regarded as a continviation of the one dealing with the genera 

 of Viverrinje which was published in the 'Proceedings' for 

 March 1915 (pp. 131-149). Its subject-matter is treated on the 

 same genei-al line^ as those therein adopted, and its main purpose 

 is to show first that the genera discussed, hitherto diagnosed 

 mainly by cranial and dental characters, may be equally well, 

 perhaps better, distinguished by the cutaneous features examined ; 

 and secondly, that these features fully justify the conclusion, 

 hinted at but not adopted by Mivart, that the Yiverrine and 

 Paradoxurine genera should be relegated to distinct subfamilies 

 of the ViverridsB. 



The gen VIS Paradoxurus, including the species referred to 

 Paguma and Macrogalidia, ranges from India and Ceylon to the 

 Philippine Islands and Celebes. Arctictis extends from the 

 eastern Himalayas to Borneo, and Arctogalidia, with the same 

 eastern limit, reaches Assam. Nandinia, on the contrary, is 

 restricted to the forests of tropical Africa. 



The Feet of Paradoxurus. 



Of this genvis I have examined examples of the three species 

 P . larvatus from Szechuen, P. niger from India, and P. herina2)hro- 

 ditus from Singapore. According to Gray's nomenclature, which 

 there is a tendency at the present time to revive, larvatus should 

 be referred to the genus Paguma ; but in this pap§r I propose, 



