CHARACTERS OF THE PARADOXURIN.E. 409 



being prepenial, instead of prescrotal as in Paradoxurus and 

 others *. 



However that may be, the gland of the female Arctogcdidia 

 differs from that of Faradoxiirus in several respects. In Arcto- 

 gcdidia the principal secreting ai-ea is the depression in front of 

 the clitoris, encircled by an upstanding flap of thin skin naked 

 without and within and appai-ently not specially glandular. 

 The right and left portions of this flap form two labia confluent 

 in front and capable of being folded over the glandular depres- 

 sion, thus meeting in front of the clitoris and vulva. Between 

 the vulva and the anus there is a tolerably long hairy perineal 

 area. In the female Paradoxurus the labia are thick, glandular, 

 and haiiy externally. They are not confluent in front but 

 arise separately at the sides of the clitoris," pass backwards 

 on each side of the vulva and extend some distance behind 

 it, almost reaching the anal area, being separated therefrom 

 by a very short hairy non-glandular area. When folded over 

 they meet in the middle line behind the clitoris. 



The Perfwme-gland and Exte<i'nal Genitalia of Nandinia. 



The gland of the male was described by Flower (P.Z. S. 1872, 

 p. 684) as follows : — " Nandinia resembles many of its allies in 

 possessing a ... cutaneous scent-gland in the form of a longitu- 

 dinal median depression an inch in length, with tumid naked 

 margins and looking very like a vulva, situated in the pubic 

 region immediately in front of the short. Conical reti-overted 

 hairy prepuce." This descrij)tion is perfectly cori'ect, but it 

 unfortunately suggests similarity in position between this gland 

 and that of the typical Paradoxurines. 



Ten years later Mivart (P. Z. S. 1882, p. 170, note) made a 

 similar mistake when he described the gland as " a bald patch, 

 no doubt glandular^ in the situation of the prescrotal glandular 

 structure of Genetta" It is quite true that the gland in Nandinia 

 is prescrotal, but it does not lie between the scrotum and the 

 penis as in Genetta^ but in front of the penis as Flower stated. 



In 1900 Miss Oarlsson (Zool. Jahrb; Syst. xiii. pp. 509-528) 

 figured and described the gland of the female as a narrow area 

 of naked skin situated in front of the vulva and ending anteriorly 

 in a distinct pouch. This had been previously detected by 

 Noack, who regarded the pouch as the functional homologue of 

 the marsupial pouch (Zool. Garten, Frankfurt, xxvii, p. 79, 1886). 



In a half-grown female Nandinia I found the glandular area 

 exactly as described by Miss Oarlsson, except that it was separated 

 from the vulva by a narrow area covered with hair and had 

 only an indistinct pouch at its anterior end. It consisbed of an 

 elliptical patch of pale naked skin, with the anterior rim slightly 



* If this proves tO be so; a good case miglit b6 irlade out fcir severins Nandinict, 

 and Arctogalidia from the Paradoxuriilaj as a separate subfamily Nandiniinse. 



