Il8 CALIFORNIA ACADEMY OF SCIENCES. [Proc. 3D Ser. 



nuclei. The latter are sharply defined, relatively large, 

 and number from three to seven in each cell, or group, if 

 they be considered syncytia. In some cases these cells are 

 of perfectly sharp contour, but in other examples the anterior 

 extremities are drawn out to form a stem. Whether this is 

 the commencement of a duct has not been decided, since 

 it rapidly vanishes into the neighboring connective tissue 

 and yolk-glands. 



The penis is attached to the outer extremity of a two- 

 layered sac which is invariably filled with the secretion from 

 the prostate glands. Goto recognized this latter substance, 

 and states that "what von Linstow (1889) calls the ' Sa- 

 menblase ' in Phylline hindorffii, and P. J. van Beneden 

 (1861) the ' vesicule seminale,' situated outside the penis 

 in E. kippoglossu, is undoubtedly the prostate gland." 

 Judging from E. squaniiila, this latter statement is ques- 

 tionable. These variously named vesicles are in all 

 probability reservoirs for storing up the prostate secretion, 

 but the latter is developed in cells located some distance 

 from this point. 



In living specimens the secretion of the prostate glands 

 in the reservoirs is snowy white, and appears in marked 

 contrast to the surrounding tissues. Two delicate cords of 

 the same character (figs. 12, 13 p') may also be seen ex- 

 tending from the innermost enlargement of the prostate 

 reservoir, and may be traced backward along the sides of 

 the ovary. Here each splits up into a number of branches 

 which pass to the sides of the testes or between them, and 

 become lost beyond this point in the body parenchyma. In 

 sections these fibers are found to be made up of a large 

 number of delicate ducts which open into the prostate 

 reservoir on its ventral surface, and on the other hand, to 

 connect with cells in the vicinity of the testes. 



In the region bounded anteriorly by the testes, posteri- 

 orly by the margin of the body and the pedicel, and later- 

 ally by the inner pair of longitudinal nerve cords, are a 

 large number of gland cells. Many of these communicate 

 with the posterior sucker (fig. 12), while others connect with 



