1910.] 



CUTANEOUS SCENT-GLANDS OF IlUMIxXANTS. 847 



e' . I'cdal gland!- a deep cleft with a loiii;- aiicrturc on 

 the front of the pastern, the web e\ton<lin^'' f 'iw avds 

 as a close fold of integument from the lu-els towards 

 the front of the hoof and separating the cavity of 

 the gland from the space between the hoofs. 



Antilopin.tj (p. 887). 

 c'. Pedal glands aborted or noarlj' so on the hind fiot. 



Blti3A.i,in2i (p. 900). 



i' . Never a deep iiiterdigital cleft opening by a long orifice, with 

 sharply defined upper margin, on the front of the pastern ; 

 pedal glands when present and well developed somewhat 

 flask-shaped with a constricted orifice opening on the front 

 of the pastern above the hoofs (exc. Knpicapra), never 

 opening deep in the interdigital cleft above the heels. 



f. A depression on the lower half of the pastern in front caused 

 by the gradual downward and backward descent of the 

 skin of the middle line towards the heels, where it meet.s 

 the posterior skin to form the heel-tie, allowing the 

 hoofs to be widely or moderately widely separated 

 apically ; only rarely (in Rujiicapra) does the closely 

 folded integument extend forwards from the heels and 

 shut off the interdigital cleft from the space between 

 the hoofs ; pedal gland, when specialised, folded upon 

 itself and opening by a small orifice on the front of 

 the pastern high above the hoofs. 



RupicAPEiN^ * (p. 848) and Capein^: (p. 85( 



f. At most a shallow depression on the front of the pastern ; 

 the hoofs joined together almost throughout their basal 

 extent by an interungual band of integaiment extending 

 forwards from the heels, where it meets at an obtuse or 

 right angle the skin forming the posterior wall of the 

 pastern ; it similarly meets that of the anterior wall of 

 the pastern, which forms a continuous surface in the 

 middle line, except where its continuity is sometimes 

 broken for a short distance by the orifice of the pedal gland, 

 which, \vhen present, opens close to the hoofs ; thus, 

 except for, the glandular infolding, the anterior and pos- 

 terior walls of the pastern are widelj' separated from each 

 other in the middle line. 



ff. Hoofs truncated apically, the heels raised high above the 

 ground during progression ; (interungual integument 

 hairy throughout ; a valvular preorbital gland ; muffle 

 large ; no pedal glands) Ojreoteaginje (p. 885). 



ff'. Hoofs pointed, the entire sole and heel in contact with 

 the ground during progression. 



//. Nasal cavities considerably inflated ; skin of the heel- 

 tie posteriorly folded upon itself in correlation with 

 the backward extension of the enlarged heels ; (no 

 preorbital or pedal glands ; a pair of very large 

 inguinal glands with orifice remote from the 

 mamniip) Paxtholopin^ (p. 898). 



1i'. Nasal cavities not specially inflated ; skin of heel-tie 

 not folded, heels moderate in size. 



i. A pair of large metatarsal glands on the hind legs ; 

 no false hoofs ; interungual integument continu- 

 ously hairy ; [no preorbital or pedal glands]. 



^PYCEEiir^ (p. 918). 



* This section should, perhaps, be divided into two or more subfamilies, none of 

 the genera being closely related to one another. Saiffa should possibly come under 

 the heading / as the type of a special group Saigina^ ; but the exact' structure of 

 the feet and of their glands is unknown to me. 



