1910.] CUTANEOUS SCENT-GLANDS OF RUMINANTS. 885 



between the hoofs. This was more marked in the niiile than in 

 the female, and in the former sex the heel-tie was thinner and 

 less produced in the hind than in tlie front leg. The walls of 

 the glandular interdigital depression were scantily covered with 

 .short haix's ; but the heel-tie itself was thickly clothed with long 

 haii-s, largely overlapping the base of the hoofs on their inner 

 ;ispect. The chief difference between the feet of this species and 

 those of IiaphicerKS cavipestris lies in tlie f<ict that the interdigital 

 cleft is not overlapped above by a long integnmentnl fold, "ami 

 that the fold of skin along the l^ack of the pastern extends right 

 down to the heels, tying them together, and extends further 

 forwards towards the anterior edge of the hoofs. Fioui this 

 arrangement it comes about that the hoofs of Onrehia are much 

 less distensible than in RajMcer as ; they are also shorter: the 

 combination of these two features suggesting that Ourehia 

 habitually frequents harder ground. 



The structure of the feet in Ourehia, correlated with that of 

 the preorbital glands and the presence of well-developed tufted 

 carpal or knee glands and of a single pair of inguinal glands, 

 is strongly suggestive of affinity between that genus and Gazella. 



By the structure of the feet, Oareh'm and Raphicerus may 

 be distinguished as follows : — 



a. Orifice of glandular interdigital cleft much sliorter than the 

 cleft itself, which is partially overlapped in front and above 

 by a longish integiimental fold or web : posterior web of the 

 ])astevn not reaching the heels, which can thus bo widely 

 separated Eaphicerus. 



a'. Orifice of glandular interdigital cleft apinoxiniately as long 

 as the cleft itself, which is not to any apprecialjle extent 

 overlapped above and in front by an integuniontal fold ; 

 ])Osterior web of the pastern reaching the heels, which are 

 thus capable of being separated only to a very limited 

 extent Oarebia. 



Subfamily O R E o T K A o i n .e. 

 Genus Oreotragus H. Sm. 



Oreotragus oreotragus Zimm, (The Klipspringer.) 

 (Text-fig. 101, A, B.) 



According to Owen, this species has preorbital glands but no 

 inguinal pits. H. Smith, on the contrary, says that inguinal 

 glands are pi'esent. 



In a. foot of this species, given to me by Rowland Ward, which 

 had the bones of the pastern cut away from behind, but with the 

 skill of the interungual area and of the front of the pastern 

 entire, I could find no trace of pedal gland. The skin on the 

 middle line of the pastern in front showed no sign of depression. 

 Stopping short a little above the proximal margin of the hoof, it 

 turned sharply backwards at right angles between the hoofs to 



Proc. Zool. Soc— 1910, No. LVIIL 58 



