lUlU.J CUTAXEOI'S SCEXT-GLAXDS UF RL'MIXAXTS. 905 



The preorbitnl gland consisted of a subeirculiii- patch of black 

 skin, sparsely pitted with setal pores, whence hairs arose. In the 

 young male the gland was as large as a hazel-nut, but markedly 

 longer than deep. In section it was pink in colour and resembled 

 a cyst enveloped in the thickened skin. It was not lodged in 

 any <lepression of the skull, but merely rested on the bone of the 

 face. The hairs which appeared vipon its surface were seen to be 

 rooted deep in the gland antl to reach the surface through the 

 pores above tlescribed. No secretion could be squeezed from the 

 gland. This was perhaps due to the fact that the animal was 

 immature and had been ailing for some time before death. It 

 seems probable that the secretion naturally exudes through the 

 setal pores. Nevertheless, I have never seen the preorljital gland 

 of a living Gnu, either of this species or of G. taurinus, show 

 any sign of secretion or even of moisture. I did not cut the 

 gland of the adult female, since the specimen was I'equired for 

 preservation. 



Pedal glands, as in Bahcdis, are well developed only on the 

 front feet and appear to be absent on the hind feet. The 

 feet and the glands have essentially the same structure as in 

 Biihalis caama. In the adult female, however, the cavity of 

 the gland differed in some interesting particulars from that of 

 B. caaina above described. The posterior wall of the cavity 

 was not folded so closely upon the skin of the interinigual web 

 and of the back of the pasteiii, but ran obliquely upwards and 

 backwards, then turned forwards at a right angle, then down- 

 wards and forwards at a right angle closely in contact with the 

 skin of the front of the pastern, which it joined at the upper end 

 of the orifice at a very acute angle, forming a close fold. The 

 anterior upper web thus constituted extended nearly as far 

 downwards as the edge of the interungual web, leaving a much 

 shorter orifice for the gland than in Bubalis caama. In the 

 latter the gland was rhomboidal in section, whereas in C. gnu': t 

 was suboblong. It Avas thickly clothed with long haii-s, pro- 

 jecting like a fringe or tuft from the orifice. The roots of all 

 the hairs were covei-ed and stuck together with blackish secretion 

 smelling exactly like sour milk or cream-cheese. In the young 

 male the cavity of the gland was neai-h' intermediate in shape 

 between that of the adult and that of B. caama, the slit-like 

 orifice being much longer than in the adult owing to the absence 

 of the anterior wall, the antei'ior web or fold stopping short 

 much higher up the pastern above the hoofs. In section the 

 upper portion of the cavity resembled that of B. caama, while its 

 lower portion was like that of the adult C. gnu. 



On the hind feet there was only a very shallow hair-lined 

 depression, recalling that of Bubalis caama. 



The front and hind feet of another adult example, kindly given 

 to me by Rowland Ward, exactly resembled the one above 

 described. 



