1910. 1 CUTANEOUS SCENT-GLANDS OF RUMINANTS. 907 



group. Moreover, it seems to lue that the size and structure of 

 the preorbital gland and of the glands in the front feet, coupled 

 with the hairiness of the rhinaiium and the presence of a single 

 })air of mamuiEe, suggest tolerably close kinship between Dama- 

 liscas, the least-specialised naember of the Bubalinse, and the 

 Antilopinse. At all events, I think it may be claimed with some 

 assurance that the relationshij^ indicated is closer than that 

 between yEjjyceros and the Antilopinas. 



Subfamily R Y G i N ^. 



1 prefer to use the term Oiygiupe for the group called Hippo- 

 traginse by most modern authors, because the generic name 

 llippotragus will soon, probably, no longer be admitted as a valid 

 generic designation, being antedated both by Ozanna and 

 Egoceros. 



Genus Oryx Blainv. 



According to Owen the following species of this genus have 

 neither pi-eorbital nor inguinal glands : 0. gazeUa{= A. oryx) and 

 0. algazel {—A. leucoryx). 



Ogilby also says that these two species have neither preorbital 

 nor inguinal glands, but states that the pedal glands are large 

 and that there are two pairs of mammse. 



Oryx beisa Riipp. (The Beisa.) 

 (Text-fig. 109, C.) 



In the dried skin of a young male example of this species, for 

 which I am indebted to Mr. Gerrard, I could find no trace of 

 preorbital glands nor of inguinal glands, but two pairs of mammse 

 were present. 



Pedal glands, approximately similar in structure, were present 

 on all four feet. They opened by an elongated, somewhat lanceo- 

 late interdigital cleft on the front of the pastern just above the 

 hoofs. This cleft led into a shallow depression, which at its 

 deepest part, close to the interungual web, passed into the 

 principal part of the gland, which consisted of a moderate-sized 

 sac extending for a short distance upwards and backwai'ds almost 

 as far as the posterior integument of the pastern. Describing 

 the gland another way, it might be said to consist of a deep but 

 i-ather short interdigital depression subdi\T.ded into an anterior 

 and posterior part by a ridge of integument projecting from its 

 upper wall, thus constricting its lumen. 



The floor and posterior wall of this depression were formed by 

 the closely infolded layer of the interuugual web. Its opposite 

 walls were formed by the integument of the front of the pastern, 

 which sloped obliquely downwai-ds and forwards to the aforesaid 

 ridge, then obliquely upwards and backwards. The entire inter- 

 digital depression on both sides of the constriction was clothed 

 with longish white hairs clogged with secretion. Inferioi-ly, just 

 below the orifice, the hah's were longer and overlapped the 



