1910.] CUTANEOUS SCENT-GLANDS OF RUMINANTS. 911 



The feet were very different from those of Oryx and Hipjio- 

 tragus, being low, flat, long-heeled, and bi'oad at the point in 

 adaptation to a soil of soft sand. In section, the integument 

 of the posterior surface of the pastern was seen to be folded just 

 above the heels like that of Pantliolops. From that point it 

 passed forwards for some distance parallel with the sole of the 

 foot before turning abruptly upwards towards the front of 

 the pastei'n. Here it passed into a somewhat conical excrescence 

 or eminence, covered with short hair and defined behind and 

 laterally by a semicircular infolding or depression of the skin. 

 Behind this excrescence, the integument of the anterior surface 

 of the pastern was continued straight up this area without sigir 

 of insinking and was covered with normal longish hairs. The 

 latter inferiorly encircled the excrescence, which jutted up like 

 an island from their midst. I believe the excrescence to be 

 glandular. The structure of the two feet was the same, except 

 that the excrescence was smaller on the foot which I judged to 

 belong to the hind leg. 



Examination of the feet of a very young example now living in 

 the Gardens revealed no signs of the excrescence, but there was a 

 distinct orifice, representing, I suppose, the infolding of the skin 

 defining the excrescence in the adult. This orifice was in almost 

 the same position as the orifice of the gland in Oryx, and its 

 glandular nature was attested by the staining yellow of the white 

 hairs x'ound it. 



Addax has a marked patch of longer haii's below the eye 

 on each side of the face. This probably overlies a glandular 

 thickening of the skin, such as is present in Hiiypotragua. I have, 

 however, no dii'ect evidence of the fact. 



From the structure of the feet Addax would seem to be less 

 closely allied to Oryx and Hippotragus than these two are to one 

 another. But more and better material of the genus must be 

 examined before any satisfactory conclusion as to its true 

 systematic position can be arrived at. 



Subfamily CERVicAPRiNiE. 



Genus Pelea Gray. 



Pelea capreolus Bechst. (The Vaal Rhebok.) 



(Text-fig. 111.) 



According to Owen, this species possesses inguinal glands, but 

 has no preorbital glands. 



The absence of preorbital glands has been affirmed by sub- 

 sequ.ent authors, and coiTectly so, judging from the one specimen 

 of this species I have examined, a female brought b}- Dr. Lyster 

 Jameson from S. Africa, which lived two years in the Gardens. 

 This specimen, however, had no trace of inguinal glands. Pedal 

 glands were well developed on all four feet. They opened by a 



