1910.] CUTANEOUS SCENT -GLANDS OF RUMIXANTS. 921 



Although Gray long ago made the genus jEjyyceros the type of 

 a special family, it lias of late years found a place amongst the 

 Antilopinte. Apart, however, from the hairiness of the rhinaiium 

 and of the interungual integument of the feet, it has very few 

 '' Antilopine " features, in the strict sense of the word. This 

 latter feature, moreover, is as much Neotragine as Antilopine. 

 In other respects, the feet may be described as Tragelaphine or 

 Cervicaprine, except for the disappearance of the false hoofs and 

 the presence of the metatarsal glandular mats of hair. The horns 

 moreover, are as much Cei'vicaprine as Antilopine in character, 

 and the absence of the preorbital gland is another feature ex- 

 cluding the genus from the Antilopina?. 



Hence, taking into consideration the sum total of its characters, 

 the peeuliaiities of jEjiyceros seem to me to be best expressed by 

 setting it in a subfamily apart from all others. 



Subfamily Tragelaphine, 



Genus Tetraceros Leach, 



Tetraceros quadricornis Blainv. 



(The Four-horned Antelope.) 



(Text-figs. 116, A, B; 117, 118, 119, A, B, C.) 



Hodgson described the preorbital glands of the genus Tetraceros 

 as linear and longitudinal, and stated that inguinal glands are 

 absent and pedal glands absent or present only on the hind feet. 

 Blanford repeated this account with the additional information 

 that the pedal glands are confined to the hind feet. 



I have seen three examples of this species, a still-born young 

 one and an adult male and female. In all the preorbital gland 

 was well developed and valvular, consisting of a longitudinal 

 pocket, the upper rim of which was produced into a large over- 

 lapping flap, forming a longitudinal slit where its free margin 

 came into contact with the skin below it. When the flap was 

 lifted a blackish nearly naked area was exposed ; the posterior or 

 ocular end of this area was the deepest portion of the invagina- 

 tion, which from that point forwai'ds became gradually shallower. 

 The gland itself was thick and pale in colour. In the adult male 

 it was much largei' than in the adult female, and in the summer 

 dischai'ged a quantity of soft cheesy-yellow secretion. The odour 

 of this was not noted after death, the specimen having been pre- 

 served in alcohol ; but in the fresh female specimen the drops of 

 clear fluiil that were squeezed from the gland had a faint aromatic; 

 scent. 



There were two pairs of teats, but no trace of inguinal glands. 



The feet were constructed as in the genus Tragelaphus, the 

 hoofs being short and luit little distensible, owing to the extension 

 of the interungual integument from the heels forwards for some 

 distance before curving backwards to join that of the front of the 



