946 MR. R. I. PococK ox TUE [June 14^ 



iuul davaiiceli. Apart from the greater length of the false hoofs, 

 the feet of Elaphuras seem to ditter fi-om those of the species 

 just citeil in having the integumental groove on their upper or 

 anterior surfaces covered with short hair like that of the i-est 

 of the foot, instead of being naked at the bottom. 



0})inions of authors as to the systematic position of this Deer 

 are di\ided in accordance with the value attached to pai- 

 ticular chai'acters. Those, like (Gordon Cameron and Lydekker,. 

 who rely mainly upon the antlers, place Elapharus with the 

 American Deer; Bi'ooke, on the contrary, (le})ending largely upon 

 the structure of the lateral metacai-pals, maintained that Ela- 

 plmrns was an aberrant type of the group to which C. dcqyhuSy 

 dama, miicolor, and others belong. 



The structure of the interdigital area of the pastern — that is to 

 say, the absence of the glandular pouch and depi'ession on the 

 hind leg, and the smoothness of the interungual integument — and 

 the high position of the metatarsal gland completely bear out 

 Sir Victor Brooke's opinion as to the affiliation of Elapharus with 

 the Old World Deer, especially the Oriental types (cf. hifra., 

 pp. 96?-970). 



In opposition to ]\[r. Cameron's vieAv that KlapJinras is not 

 related to the Old World Deer, the Marquis of Hamilton (■ Field/ 

 July 1910, p. 199) pointed out that hybrids between Elaphunis 

 and CervKS have been bred in Paris and that one of these hybrids 

 formed fei-tile unions with both parent stocks at Woburn. This 

 is an important piece of evidence in favour of relationship 

 between the two genera. 



Genus Rusa Ham. Smith. 



RusA uxicoLOR Bechst. (The Snmbar.) 

 (Text-tig. 131.) 



According to Ogilby, this species has distinct and movable 

 preorbital glands and large pedal glands, the same epithets being 

 applied to them as to those of V. elaphus, suggesting identity of 

 development and structui^e. Hodgson supplements this descrip- 

 tion by saying that the preorbit;d glands are very large and 

 completely reversile, that the pedal glands are large and present 

 on all fovir feet, and that the metatarsal gland is posterior and 

 external. According to Blanford, however, Hodgson says the 

 interdigital glands are wanting. I have seen one old male 

 example of this species. 



The preorbital gland was a large and deep, naked invagination 

 half filled with a (juantity of waxy secretion with a strong but 

 indescribable odour. 1'he integument at the bottom of the pouch 

 was thin and apparently not glandular; but I'ound the orifice, 

 especially anteriorly, it was much thickened, and it was here 

 apj)arcntly that the secreting area was concentrated. 



The supraorbital gland, mentioned by Weber as present ia 



