HERMAPHRODITISM IN A LIZARD. 225 



and situated in the broad ligament parallel to but separate from 

 the testes. They were not attached, as is normally the case, to 

 the inner border of the testes, and vasa efFerentia were conse- 

 quently absent. Posterioi-ly, the epididymes passed into the 

 vasa deferentia, and these, having united with the ureters, opened 

 into the posterior division of the cloaca by the urogenital papillas. 



The two kidneys were apparently normal, each consisting of 

 an anterior and posterior lobe (in the figure, only the right 

 kidney, U.K., is indicated by reference letters). 



The copulatory organs (li. Co^J., L. Cop.) were present as in 

 the normal male lizard. 



The oviducts {R.M.D., L.M.D.) were developed for about 

 a third of their lengths. Each opened into the body-cavity by a 

 well-developed funnel (only the left, L.FL, is indicated by 

 reference letters in the figure), and behind that was continued 

 into the duct, the plaited glandular walls of which are seen in 

 the outer border of the broad ligament (B.Lg.). The right 

 oviduct attained the greater complexity as in the normal female. 



Passing down from the posterior tip of the ccelomic funnel on 

 each side, on the extreme outer border of the broad ligament 

 was a narrow but well defined ribbon-like muscular band which 

 continued right back on each side to the cloaca.. Similar bands, 

 which were at first taken for the oviducts, were referred to by 

 Howes (5) as the round ligaments, and he thus describes them 

 in his specimen : " From this (the oviducal aperture) there passed 

 back a ribbon-shaped muscular band which skirted the free edge 

 of the broad ligament, remindful of the round ligament of 

 mammalian anatomy. This structure was wholly absent on the 

 side destitute of an oviducal vestige, as indeed it is in the normal 

 male. Its development is correlative with that of the oviduct." 



b. Histology. 



Transverse sections were made of the gonads and stained with 

 hasmatoxylin and eosin. The main body of each gonad was found 

 to consist of normal testicular tissue, i. e. seminiferovis tubules 

 with an interstitial sti^oma, the lining cells being in active 

 mitotic division (text-fig. 2, s.t.). 



The stalked outgrowths were found to consist of ovarian tissue, 

 the bulb-like extremities containing large and fully-grown ova. 

 A section of the gonad through the nuclear plane of the most 

 anterior ovum is somewhat diagrammatically represented in 

 text-fig. 2. 



The large yolk-laden ovum {ov.) is, it will be seen, surrounded 

 by the relatively thin follicular epithelium {foil, ep.), outside 

 this is a fibrous layer continuous with that sheathing the 

 testicular portion of the gonad. In the stalk region all the 

 normal histological elements of the ovary are represented. 

 Outside is a layer of cubical epithelial cells continuous with 

 the peritoneal epithelium, while the main mass of the stalk is 

 formed of a loose stroma of connective tissue, contained in which 



