174 MR. F. E. BEDDARD ON THE [| Feb. 16, 
For example, in the brain of Madoqua phillipsi, which will be 
shortly described in the present communication, the splenial 
suleus is continuous anteriorly with the crucial suleus and does 
not communicate with the coronal sulcus, which sends a branch 
osteriorly to the middle line. Precisely the same arrangement 
occurs in the Sheep, as figured by Elhot Smith. On the other 
hand, in the Pig tribe, as is well known, and as will be also 
pointed out in the present communication to the Society, the 
splenial fissure 1s continued into the coronal. In the Babyrussa 
there would seem to be a continuation of the splenial imto both 
these fissures. The crucial fissure and the fissure lying behind it, 
which represents the inner and posterior limb of the sulcus coro- 
nalis, are as plainly shown in Antilocapra as in Madoqua (cf. text- 
figs. 11 & 16) and in other genera *. The crucial fissure on one side 
the right) joins the coronal, and on the other though continuous 
with the coronal it does not reach the middle line of the brain. 
The coronal behind is also asymmetrical, but in a different way ; 
on the right the fissure is separated by a bridging convolution 
from the junction of the coronal and the branch of the supra- 
sylvian. On the left there is complete continuity. 
Tt is this coronal fissure which is continuous with the splenial, 
and not the crucial as in Ovis. The lateral fissure proper, which 
T take to be the middle one of three lateral fissures, is remarkable 
for the fact that at the end of its first third (about) there is a 
strong branch at right angles on the outside, and also some indi- 
cations of a similar branch at the same spot on the inside, the two 
together forming a cross branch. The ecto- and entolateral sulei 
run forwards to nearly as far as this point. The entolateral 
fissure, it may be observed, does not nearly reach the middle line. 
The suprasylvian fissure is absolutely continuous with the coronal, 
as in Cervus. It bifurcates posteriorly in the occipital region 
intoaY. There are two marked descending furrows, one in front 
of and one behind the Sylvian. The genwal fissure is not con- 
tinuous with the splenial (intercalary). 
In addition to the brain of the adult Prongbuck just described, 
IT have had the opportunity of studying two fetal brains 
extracted from the mother, but of nearly if not quite full-time. 
Unfortunately they were both rather damaged at the time, but 
T have been able satisfactorily to compare certain of the sulei with 
those of the large brain. The strong branch of the lateral sulcus 
running at right angles to it, which seems to me to be so charac- 
teristic of Antilocapra, is quite obvious in both of these brains 
and on both sides of the brains. I have mentioned particularly 
the fact that the entolateral suleus does not reach the middle line 
of the brain. In one of the feetal brains the relationship was the 
same; inthe other, however, the entolateral sulcus of the left side 
only did dip over the mesial edge of its hemisphere. As to the 
continuity of the coronal with the splenial sulcus I can only 
* BE. g. Cervus, Elliot Smith, loc. cit. p. 329, fig. 190. 
+ Holl, Arch. f. Anat. Phys. pl. xviii. fig. 10, & Elliot Smith, Zoc. ci¢. fig. 191, p. 329. 
