188 MR. F. E. BEDDARD ON THE [ Feb. 16, 
5) Some Notes unon the Anatomy of Madoqua phillipsi. 
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A dissection of an example of this small Antelope enables me 
to make a contribution to the knowledge of the structure of the 
Cavicorn Ungulates, for I believe that we have at present no 
information as to the anatomy of this pentigmbn genus of 
Antelopes, beyond the notes contained in Lénnberg’s paper already 
referred to. 
The example which I examined was a female. The usual 
four teats were present and also as usual the anterior pair were 
much further away from each other than the pair lying behind 
them. There was no trace that I could see of the inguinal 
glands, and, indeed, the inguinal region, instead of being bare, 
was particularly well covered with long hair. I have notes 
upon some of the organs, and I naturally paid special attention 
to those structures which are known to vary in these animals. 
As to the Vascular System I found that the heart shows the 
“‘moderator band” so frequently found in running animals— 
birds as well as mammals*. It was a considerable fleshy 
column connecting together the free and septal walls of the 
right ventricle. The postcaval vein was typical in its arrange- 
ment, 7.e. the vein lay postrenally to the right of the aorta. 
The azygos had the following structure :— 
The general plan is like that of other Artiodactyles. That is 
to say, “the ri ight azygos is much less important than the left. 
The latter opens moreover, as is usual, directly into the heart, 
but it consists of two portions, an anterior and a posterior. 
They unite just opposite to the transversely running vessel 
which is formed by their union and debouches into the heart. 
There is thus, as I may point out, a remnant of the left anterior 
cardinal, which I presume the anterior branch of the left azygos 
to represent. This fact in the structure of the azygos of 
Madoqua is furthermore of some systematic interest. For, as 1 
have discovered tT in Rhaphicerus and Ourebia, the arrangement 
of the veins bringing back blood from the dorsal parietes on the 
left is quite like that which has just been described in J/adoqua. 
Now these three genera have been placed in the same subfamily 
Neotragine. This additional structural likeness justifies still 
further this placing. The right azygos of J/adoqua is but feebly 
developed and brings back blood from only two or three inter- 
costal spaces. 
In the intestinal tract of Antelopes it is the colon which 
shows the principal differences. I have therefore made careful 
notes upon certain points in the structure of that part of the gut 
in Madoqua phillipsi. The cecum is long and like that of 
other Antelopes in its blunt ending without any diminution 
of calibre. The iewm enters it at an acute angle and is attached 
to it hy mesentery which extends a long way towards the tip of 
* To which particular attention was called by the late Prof. Rolleston. 
+ “On the Azygos Vein in the Mammalia,” P. Z.S. 1907, pp. 184, 185. 
