190 MR. F. E. BEDDARD ON THE [ Feb. 16, 
Dr. Linnberg has specially devoted himself to the intestinal 
tract (and other features in the anatomy) of the Artiodactyles ”. 
As these animals are comparatively little known in the anatomy 
of their soft parts, Ido not hesitate to set down some further 
facts as a contribution to our gradually increasing knowledge of 
the group, which may prove to be distinctive not of the particular 
species now under consideration but of the group generally. In 
Madoqua, as in Cephalophus abyssinicus, the end of the duodenal 
loop is not fixed by the ligamentum cavo-duodenale to the post- 
caval vein or to the mesocolon, but to the ventral surface of the 
colon itself. 
Some of the recto-duodenal ligamentum strays over on to the 
adjacent right kidney. I could find no definite hepato-renal 
ligament. But the right extremity of the liver was attached to 
the parietes by a well-marked ligament. I may remark also that 
in Cephalophus abyssinicus the duodenum was attached to the 
rectum itself in precisely the same way by a ligament, and that 
there was also the same attachment to the right kidney. The 
left kidney was situated a long way below the right kidney, 
there being a considerable distance between the posterior end of 
the right kidney and the anterior end of the left kidney. The 
spleen of Madoquwa had the more or less circular contour of this 
‘“oland” in some other Ruminants. It was as firmly fixed to the 
stomach throughout as in other Ruminants. There was in fact 
no free gastrosplenic omentum. 
The Brain of Madoqua phillipsi is not known, so far as I am 
aware, and will serve to illustrate the brain characteristics of a 
small Antelope with the suici in a simple condition. It may be 
compared with that of Antiocapra, which I also figure in the 
present communication. The general outline seems to me to be 
remarkable for the square outline and termination anteriorly of 
the portion of the brain lying in front of the Sylvian fissure. 
This form of brain is still more marked in the Pronghorn, which 
T have dealt with on a previous page. Secondly, as will be seen 
in the drawing of the dorsal surface submitted herewith (text- 
fig. 16), the corpora quadrigemina are partially visible, which is at 
least unusual among the Ungulata. The third obvious peculiarity, 
also to be seen in the same dorsal view, is the exposure of the 
splenial suleus which runs on each side a curved course at some 
little distance from the median fissure of the brain. The exposure 
of this fissure is not unknown in other Artiodactyles—it occurs, 
for instance, in Vragulust, Dorcatheriwm, Moschus, Cervulus— 
but it must not be confused with an entolateral sulcus, such as 
occurs in Cervus damat, and which at first sight might be 
confused with it. Its continuation with the crucial anteriorly 
* Nova Acta Reg. Soc. Upsala, 1903; K. Vet.-Ak. Handl. 1901; P.Z.S. 1900, 
p. 142; and in Zoologiska Studier tillagnade Professor T. Tullberg, 1907, p. 237. 
+ See Cat. Physiol. Series, Roy. Coll. Surgeons, Vol. 11. 2nd ed., 1902, p. 323, 
p. 324, p. 325 fig. 187, p. 327. 
+ Ibid. p. 329, fig. 191. 
