1909.] ANATOMY OF CERTAIN UNGULATA. 191 
shows that in Wadoqua the fissure in question is really a splenial. 
Dr. Elliot Smith has remarked * that “the position of this sulcus 
in the Ungulata seems to be determined largely by the size of 
the hemispheres,” it being visible above in Zragulus and not in 
Cervus. So far as Madoqua is concerned this generalisation 
would seem to be true of the Cavicornia as well as of the other 
Ruminants. The brain of the Sheep has been so often described 
and figured 7 that it may be well regarded as a convenient basis 
of comparison for other hollow-horned Ruminants. There are in 
Madoqua, as in Ovis, three lateral fissures of which I identify 
the middle one and the longest as the lateral, the other two 
being ecto- and ento-lateral respectively. The lateral bends 
outwards anteriorly, but does not end in any other Sssure. 
Text-fig. 16. 
Brain of Madoqua phillipsi, dorsal view. Natural size. 
a. Coronal suleus. _c. Inner branch of the same. d. Suprasylvian sulcus. 
e. Lateral sulcus. f Entolateral sulcus. g. Splenial sulcus. 
The suprasylvian fissure is the same on the two sides of the 
body and consists of precisely the same elements in the Sheep, 
excepting that I did not find the fissure lettered by Holl £ “rdss” 
and represented in the drawing given by Elliot Smith not far 
behind the Sylvian fissure, but not lettered. The suprasylvian 
fissure, however, bifurcates posteriorly in a V-shaped fashion as 
indicated by Holl in Ovis; but in Madoqua the two limbs are 
longer, the downwards directed limb running parallel with the 
suleus obliquus. The anteriorly situated “inferior” branch of 
the suprasylvian joins the sulcus coronalis. 
%* Tbid. p. 326. ost 
+ Most recently by Elliot Smith (loc. cit. p. 340, figs. 194, 195, 196) and Holl, 
Arch. f. Anat. 1900, pl. xvii. figs. 7, 8, 9. 
{ Loc. cit. pl. xvi. fig. 7. 
