1909. | ANATOMY OF CERTAIN UNGULATA. 193 
specimens of Sus cristates and Dicotyles tajagu, of both of which 
brains exist in the collection at the Prosectorium. 
The coronal sulci are much more obliquely set than in either 
Sus or Dicotyles, especially more so than in Dicotyles, where the 
furrows in question, save just where they dip into the middle 
line, are straight and parallel with the long axis of the brain. 
The brain of Babyrussa in fact agrees in this particular with 
that of Phacocherus ethiopicus figured by Elliot Smith *. Not 
so, however, in their proportionate length; for while in Phaco- 
cherus—judging from the figure already referred to—the coronal 
suture of each side is very nearly half the length of the brain, 
the length of each of these sulci in Babyrussa is 37 mm., measured 
from the anterior end of the brain to the point where they meet 
in the middle line of the brain; the rest of the brain measures 
about 55mm. ‘These measurements are taken along the curvature 
of the brain, and are therefore in excess of that just given of the 
length of the hemispheres. 
In Dicotyles tajacu the proportions are reversed, the anterior 
part of the brain to the posterior end of the coronary sulci is 
actually longer than the rest of the hemispheres. In Svs, 
however, the proportions would appear to be very much as in 
Babyrussa, and probably by making allowances for the curvature 
not measurable in the figure quoted, Phacochwrus is not very 
different. Elliot Smith agrees with Garrod? in regarding as 
distinctive of the Pig tribe the blending of the coronal and 
interealary (or splenial) sulci as compared with other Artio- 
dactyles. 
The intercalary or splenial sulcus in Babyrussa (which obviously 
suggests—if it be not comparable to—the calloso-marginal of 
Apes) is continuous with the genwal sulcus in front, as in Sus 
and Dicotyles. It is also continuous with the coronal, bifurcating, 
in fact, as it were anteriorly to form the coronal and genual. In 
this Babyrussa of course agrees with other Pigs, as has been 
already mentioned. It does not, however, agree with an example 
of Dicotyles tajacu which I have in my possession. I am not 
quite able to understand precisely what the late Prof. Garrod 
meant when he wrote concerning the splenial fissure of Dicoty/es 
as follows £:—“ There is one upward branch of the splenial fissure 
which joins § the fissura coronalis, and is not a continuation of it, 
as in Sus.” Dr. Elliot Smith in dealing with the brain of the 
same species repeats Garrod’s word “joined.” I find in the only 
brain which I have exainined no communication whatever between 
“the cingular are of fused calcarine, intercalary and genual sulci” 
and the coronal sulcus on either side of the body. <A bridging 
convolution occupies the place where this fissure would otherwise 
lie. But if the coronal sulcus were continued back to join the 
* Cat. Physiol. Series, Roy. Coll. Surgeons, vol. 11. 2nd ed. 1902, p. 316, fig. 182. 
+ “On the Brain of Hippopotamus,” Trans. Zool. Soc. x1. 1880, p. 13. 
i Hoc. cit. p. 14. 
§ The italics are Mr. Garrod’s. 
PRoceZo0oLy SOU.) 909s Non xelule 13 
