810 MB. F. E. BEDDARD ON THE [NoV, 12, 



it is very large and clijSfers in its form fi'om that of some other 

 genera of Yiverrines. In Suricata tetradactyla Sir Richard Owen 

 remai'ked * that " the pancreas has a singular form. A thick 

 transverse portion extends from the spleen behind the stomach 

 to the pylorus ; it then divides and forms a circle, which lies in 

 the concavity of the great curve of the dtwdenum ; sending off 

 •one or two processes in the inesoduodenwrn" This peculiar 

 pancreas was later figured by Owen f. A pancreas of this form, 

 is, however, neither universal among the Viverrines nor confined 

 to that group. Dr. Mivart's figures of the stomach and pancreas 

 of Genetta tigrina J prove the former statement, and my own 

 figures of Helictis and Galictis § prove the latter. "With regard to 

 Genetta tigrina^ I may remark that G. vulgaris has quite the same 

 form of pancreas, i. e. not forming a figure 6 as in the Suricata. 

 The pancreas of Genetta vulgaris gives off a small lobe running 

 towards the liver by the side of the bile-duct, and on the other 

 side of the bile-duct — the left — is a comjDletely detached lobe of 

 pancreas whose connection with the rest of the pancreas I found 

 it impossible to detect. I may take this opportunity of remarking 

 that in Arctictis hwiturong the pancreas, which was not described 

 by the late Prof. Garrod || in his account of the anatomy of that 

 animal, is on the whole like that of the Genet ; that is to say, it 

 is a sti-aight or rather chevron-shaped gland forming no circle 

 round the duodenal loop. In Galidictis striata the appearance of 

 this gland is quite different. For it runs round the loop of the 

 duodenum as in the Suricate ; but it does not, as is the case with 

 that animal, rejoin itself in the region of the pylorus. There is 

 also a small process of the pancreas i-unning up alongside the 

 bile-duct as in Genetta, but no detached lobe. An anangious 

 fold of membrane attaches the gastric region of the paiicreas to 

 the median mesentery as in other Oarnivora ^. It is very much 

 as I have figured it in Helictis personata, but runs up to very 

 nearly the end of the pancreas. 



§ The Postcaval Vein and its branches. 



Information upon the venous system of these Oarnivora is so 

 scanty, that no apology is needed for giving a description of such 

 facts as I have observed in Galidictis striata and in Genetta vulgaris, 

 with which I have been able to compare it. The accompanying 

 drawings (text-figs. 213, 214) show the veins in question in the two 

 Yiverrines. In both of them the branches on the two sides of 

 the body are asymmetrical ; but the asymmetiy diflers in the 

 two animals. In Galidictis (text-fig. 213) the veins are arranged 



* P. Z. S. 1830-31, p. -40. 



t Comp. Anat. vol. iii. 1868, p. 444, fig. 351. 



X p. Z. S. 1882, p. 505, fis:. 6. 



§ P. Z. S. 1905, p. 25, fig. 9 & p. 26, fig. 10. 



II "The Anatomy of the Binturong," P. Z. S. 1873, p. 196. 



% Beddard, " On Helictis personata," P. Z. S. 1905, p. 24. 



