316 ' DR. T. GOODEY ON 



doubt serve as a source of food. There does not appear to be a 

 contractile vacuole. The body measures from 8-12 jj. in length. 



Fixed and stained material shows that the protoplasm is 

 alveolar in structure, and that the nucleus is situated at the 

 anterior end of the body, as in Trichomonas and Trichoniastix and 

 similar forms. The nucleus is a vesicular structure of variable 

 size, but often of quite large dimensions. There is no central 

 karyosome, but the chromatin is disposed irregularly in two or 

 more masses. Very frequently there are two semilunar blocks 

 of chromatin situated on either side or on the anterior and 

 posterior tafcrders of the nucleus. It will be seen from the 

 iigures in what an irregular fashion the chromatin is arranged. 



The fiagella arise from blepharoplasts placed just anteriorly to 

 the nucleus. They are four in number and, as stated above, are 

 disposed in two pairs. Thej^ take their origin in four blepharo- 

 plasts, which when seen in side view (fig. 25) appear as two large 

 granules in contact with each other, but when seen in face view 

 are easily recognisable as being a group of four distinct granules 

 in intimate contact (figs. 26 & 27). The posterior pair of granules 

 is connected with the nucleus by means of two rhizoplasts, 

 which appear as one short rod in fig. 25 but are well shown 

 in the ventral view obtained in fig. 26. I made a very careful 

 examination of this region, in order to determine if rhizoplasts 

 were present connecting the anterior pair of blepharoplasts 

 with the nucleus, and I am satisfied that the pair connecting the 

 posterior blepharoplasts is the only one. 



(b) Reproduction. 



I will deal first with the multiplication of the fiagella, because 

 this always occurs prior to the division of the nucleus. The new 

 fiagella are produced by outgrowths from the body of the organism, 

 and not by splitting of the old fiagella. In this it resembles Copro- 

 7nonas, Trichomonas, and many other flagellates. The process is 

 initiated by the growth in an anteidor direction of each pair of 

 blepharoplasts : a point very difiicult to make out in many of the 

 organisms, but well shown in other cases, one of which is repre- 

 sented in fig. 27. The flagellar buds arise as delicate hair-like 

 outgrowths from the developing buds of the blepharoplasts. Each 

 pair of flagella has thus a new pair of fiagella produced imme- 

 diately anterior to it. The new ones do not stain so deeply as 

 the old fiagella, as will be seen from the figures representing 

 different stages of the division of the nucleus. In this way the 

 original posterior pair acquires a, new anterior pair, and the original 

 anterior pair grows longer and becomes the posterior pair of one of 

 the daughter-organisms, at the same time acquiring a new anterior 

 pair. The four pairs of fiagella thus produced gradually sepai'ate 

 into two sets of two pairs, which finally come to take up positions 

 at either end of the anterior face of the nucleus; but this migra- 

 tion takes place at different peiiods dining the progress of nuclear 



