1903. | NEW SPECIES OF EARTHWORMS. 219 
the spermatheca of Heliodrilus. For this reason I cannot agree 
to Michaelsen’s placing of my genus dAlvania within the genus 
Hyperiodrilus. The former has a true spermatheca, homologous 
with that of other non-Eudrilid earthworms, while the latter has 
not. ‘This also is the case with the genus or genera with which we 
are dealing now. Pareudrilus stagnalis has a spermathecal sae 
which communicates with the ccelomic sacs envolving the ovary ; 
“ Pareudrilus” papillata has not. It is possible, therefore, that we 
should revive Michaelsen’s Unyoria for the latter species. The 
stalk of the egg-sac, as might be expected, lodges the funnel of the 
oviduct, or, to speak more accurately, the greater part of the funnel. 
The exact conditions obtaining are the following. In a series of 
sections it may be seen that the narrow tube communicating with 
the ccelomic sac surrounding the proximal end of the spermathec: 
runs forwards and opens into the cavity of the xilith segment bya 
wide orifice ; its walls are therefore continuous with, and no doubt 
developed from, the septum bounding segment xiii. posteriorly, 
The upper ‘ lip” of the ostium is covered by the cubical cells of 
the oviduct, which here opens freely into the cavity of the xiiith 
segment. Further on in the series of sections, the mouth of the sac 
surrounding the spermatheca is closed, and the tube opens into the 
egg-sac through a wide tube which is entirely lined by the cells of 
the oviducal funnel ; these are, of course, perfectly continuous with 
those cells which he in the xiiith segment. The part of the 
oviducal funnel which les in the xivth segment appears to be 
divided into two, and to open by as many mouths into the huge 
egg-sac, which is so kidney-shaped as to be nearly divided into two 
sacs; I cannot pretend to an accurate description of the funnel and 
its various foldings. It is clear, however, that the conditions which 
obtain are those of the more typical EKarthworms, where the funnel 
opens partly freely into the xiith segment and is partly reflected 
so as to open within the ege-sac. Now, the simpler forms of 
Kudrilidee, such as the genus Hudriloides, are distinguished by the 
fact that the ovary is unenclosed in any sac, and that the funnel 
of the oviduct opens precisely as has been just described in Pareu- 
drilus papillata. On the other hand, in the more complicated 
forms, such as Séuhlmannia, the sacs containing the ovaries 
envelop also the oviduct-funnel and communicate with the 
spermathecal sac. The species which forms the subject of the 
present remarks is plainly intermediate between these two 
extremes; and for that reason, as I think, deserves generic 
separation from Pareudrilus. i 
§ On the Spermatophore. 
The spermatophores of the Eudrilide have not been much 
studied, and, so far as I am aware, are known only in the genera 
Stuhlmannia and Polytoreutus, in which I have myself described 
them. Many, if not most, of the individuals of the present species 
which I examined had a single spermatophore in both the 
spermathecze, 
