> Sy 
THE PISTILLATE FLOWER 7 
(TEXT FIG. 3). Whether these nuclei move about in the fluid 
mass or are carried among the spores by the movement of the 
protoplasm escaping from the tapetal cells has not been deter- 
mined, but the forms of the nuclei and their even distribution 
seem to indicate individual movement. That these free nuclei 
perform some life function is indicated by their persistence up 
to the maturity of the pollen spores. 
THE PISTILLATE FLOWER ~ 
The pistillate flowers arise as broad, contiguous protuberances 
on the lower portion of the short conical tips of the central axis 
(TEXT FIG. 4). The development of the ovary wall and the 
=. 1 Ag 
Hea Loihi ; ¢ pintillate opil bi 
only. 
Fics. 5, 7, 8, 9, 10 show in diagram, successive stages in development of ovary- 
Fic. 6 is a cross section of 7 through 
FIG. ro represents a longitudinal section of an almost mature ovary, showing 
the beii of the stigmatic hairs at S. All X 30 (nearly). 
beginning of the ovules is shown in TEXT FIGS. 5-10. A section 
of a mature ovary is shown in PLATE 2, FIG. 33. The one to six 
orthotropous ovules develop from a small basal placenta. The 
placenta represents in this case the end of the branch axis, and 
the ovules, when more than one, seem to be lateral outgrowths. 
As the ovary wall closes above the cavity the margin is broadened 
and flattened so that the lower edge closes first, leaving a funnel- 
like opening (TEXT FIG. 9). Continued growth of the wall and of 
