26 Journal of the Asiatic Society of Bengal, [January, 1909. 
secondary branches are produced by the development of main 
and accessory buds which have been shown to exist. The 
secondary branching may, however, be either monopodial or 
sympodial, and the various species of Indian cottons now under 
observation and experimental cultivation exhibit considerable 
variety in the manner of occurrence of the two types. The two 
extremes are very marked ; in the first of these the main axillary 
bud gives rise to a monopodium, while in the second, with the 
possible exception of a few lower branches, this bud gives rise 
ium.' In this latter case the secondary branches 
are composed of sections formed by the development of the 
main axillary bud, and the accessory bud occupies the leaf axil. 
Such a branch, therefore, appears at first sight to be a mono- 
podium with a single axillary bud, but can at once be distin- 
guished by the scar formed by the withered terminal bud and the 
slight angle at which the segments lie with respect to each other. 
The accessory bud may develop as well as the main bud, 
and, when this occurs, the accessory branch usually differs from 
the main branch inthis character. Thus, when the main 
i ud gives rise to a monopodium, the accessory bud 
will form a sympodium, or, if the main branch is a sympodium, 
the accessory branch will be a monopodium. 
The structure of the tertiary branches has not been followed 
in sufficient detail to justify any definite conclusions, but when 
they occur, they commonly differ from the branches from which 
ey arise—in other words, the main axillary branch of a second- 
monopodium ; while, in the case of a sympodium, the secondary 
bud (which alone remains) will give rise to a monopodium. 
On crossing a monopodial by a sympodial type, the 
offspring differ very slightly from the sympodial parent, though 
the number of monopodial secondary branches may be some- 
what increased. In the next, (F,), generation, from self-fertil 
( ; , every proportion 
of sympodial and monopodial branches occurring on a single 
stem is found. The change from the monopodial to the sym- 
podial type of secondary branch, when both occur on the same 
stem as developments of the main axilla buds, is abrupt, the 
ympodial the apical 
flowers are not concentrated into any well-defined 
inflorescence, but are developed on the sympodial branches from 
the terminal bud, and, therefore, appear to lie opposite the leaf. 
: ry branches are sympodia, these 
branches will themselves bear flowers. In types, however, in 
1 These two types will, for brevi ; 
and sympodial types respe etively. revity, be referred to as monopodial 
