DE. J. MURIB ON THE OEGANIZATION OP THE CAAING WHALE. 257 



membrane, or septal semidivision. This gives a median pouched character to the first 

 chamber, and doubtless is what Jackson mistook in his inflated specimen for a supple- 

 mentary cavity of a crescentic form opening largely into the first. The interior of the 

 fijst stomach is of a white or opaline colour, lined with an epithelial layer of membrane 

 -^ of an inch in thickness ; this ends abruptly and ring-like at the entrance into the 

 second stomach. The muscular and serous coats are moderate layers. 



The second chamber or true digestive cavity (//), as Jackson says, is a globose bag, 

 and in our female, from upper to lower border, following the curve, measured 11 inches, 

 and from left to right 8 inches. It lies partially in front and to the left side of the first 

 chamber ; and its upper posterior wall is adherent to the oesophagus by strong cellular 

 tissue. Its inferior wall approximates both to the third and fourth chambers. The 

 raucous rugse of this second stomach give its great thickness. The folds are large free 

 lappets, which radiate from the aperture of entrance towards its left wall, and in so 

 doing wind in long, parallel, crooked lines, with innumerable subsidiary shorter diagonal 

 intersecting plicae connecting them. In these interdigitations they resemble the cha- 

 racter obtained in the longer second stomach of the Porpoise. The mucous membrane 

 of this second cavity in the Deductor, as in the latter Cetacean, is of a rich florid colour, 

 and very vascular. The communicating aperture or passage leading from this the 

 second to the third chamber is a constricted narrow tube or canal, 4? inches long, 

 tunnelled between the walls of the second, fourth, and partly the thii'd stomachs. It 

 leaves the second stomach on its right inferior wall, an inch below the wide aperture 

 which connects the first and second, and it enters the third stomach above and behind. 

 There is no true sphincter at either end. 



The third gastric chamber (///) is also subglobular when distended, and occupies the 

 lower interspace between the second and fourth cavities. Its diameters or, rather, semi- 

 circular dimensions are 3^ inches longitudinally and 4 inches transversely. The walls 

 are thin, and the internal mucous coat nearly smooth and devoid of ruga, although it 

 may be noted that to the left side, and below the aperture leading to the fourth cavity, 

 there are faint traces of longitudinal plications. The round, smooth, ring-like orifice 

 between the third and fourth stomachs, is a little over half an inch in diameter. It is 

 an inch distant from the opening into the second and third ; and its direction is nearly 

 at right angles to it. 



The fourth digestive chamber (IV) is in the main long and cylindrical. Its first por- 

 tion, or left end, however, is rather wider, 2^ inches deep from the orifice, and bulges, 

 in a pouch-like manner, upward and towards the right wall of the second stomach ; 

 between which cul-de-sac and the second stomach the tunnelled canal spoken of as 

 connecting the second and third chambers runs. This sac-like part, viewed externally, 

 seems only a portion of the third stomach, but when opened is perfectly separate from it. 

 The remainder of the chamber narrows as it proceeds to the pylorus. The total length 

 of this compartment is 18 inches; following the curvature of the dilated first portion it 

 is 3-^ inches in breadth, and the remainder is about 2^ inches. Its walls are smooth, or 



