VAILLANTIA HISPIDA 17 
form haustorial appendages. In some cases when two longitu- 
dinal divisions have taken place in a single suspensor cell, a pair 
of the resulting quadrants grow out together, forming double 
haustoria (Pl. 4, Fig. 5). Again the haustorial cells may divide 
transversely ; this occurs only in the region near the embryo proper. 
These haustoria, for such I regard them, are at first pro- 
nouncedly vacuolated, but as the embryo ages the vacuolation 
disappears, and they become densely filled with cytoplasm. After 
the embryo proper begins its development, the basal suspensor 
cell elongates considerably, increasing its length by a half and be- 
comes less vacuolated. Its micropylar end becomes crushed by 
the growing endosperm, which now begins to grow into the 
micropylar part of the thickening integument, leaving the embryo 
in a more central position (Pl. 4, Figs. 9, 10). Later, the basal 
cell and even one or two of its neighboring suspensor cells become 
disorganized, but the time at which this occurs is not constant, for 
while I was not able to find these cells in the preparations from 
which Figs. 5 and 6, Pl. 4, were drawn, I did find them in another 
specimen of such an age as the one shown in Pl. 4, Fig. ۰ 
After the embryo proper has differentiated itself from the sus- 
pensor, the latter may be seen to be divided clearly into two 
regions which differ very markedly in structure. These two 
regions may be termed the micropylar and embryonal. The 
latter is made up of half a dozen disc-shaped cells which are very 
much flattened longitudinally, and show no outgrowths and are 
not vacuolated, and they remain so during the entire embryonic 
history. The cells of the micropylar region, on the other hand, 
become larger and thus increase their absorbing surface. When 
they reach their maximum development the suspensor when dis- 
sected out resembles a “bunch of grapes." These outgrowths, 
to which we have ascribed a haustorial function, are not unlike 
the suspensorial haustoria (Keimtrágerblasen, boyaux) described by 
` Treub in the Orchidaceae and by Hofmeister“ (49) and Guignard 
(2. c.) in Sutherlandia, and are analogous to the “ zuigblasen” oc- 
curring in the teak (Tectona grandis) described by Koorders (92). 
The upper part of the micropylar region, as shown above, becomes 
less important and is finally lost by absorption, so that Ihe emo 
* Die Entstehung des Embryo der Phanerogamen. Leipzig, 1849. 
