58 



the primary root shows the characteristic tuberous form, while 

 no secondary roots are yet developed- One of these is, 



* "Hypocotyl " has been used instead of the older term '' caulicle," in ac- 

 cordance with Darwin's usage (The Power of Movement in Plants, 1881, p. 5), 

 signifying the hypocotyledonous portion of the stem. 



J 



■I 



t 



Coulter, we find it as a true genus AnemoneUa^ separate from 

 Anemone and Thalictrttm. 



The plant seems to show characters intermediate between 

 those o^ Anemone and TJialictriini^ having the flower and in- 

 volucre of an Anemone and the foliage of a TJialictrnm. It 

 has umbellate peduncles like those of Anemone narcissiflora 

 and others, while the ribbed achenia ally it to Thal- 

 ictrtun ; but it does not seem, however, as though this diver- 

 gence of characters were sufficient to entitle it to rank as a 

 separate genus. 



And when the genus Anemonella, in the synopsis of the 

 genera in the revised edition of Gray's Manual, is char- 

 acterized as having a larger number of achenia than 

 Thalictriim, that is, ** four to fifteen in Anemonella^' and few 

 in Thalictri0n, it does not correspond to what is said in the 

 generic diagnosis of TJialictrMin, which is also described as 

 having *'four to fifteen achenes." 



In regard to the structure of the roots our plant shows a 

 certain peculiarity, which removes it at once from Anemone 

 and Thdlictrum. The roots are fusiform and tuberous, cor- 

 responding to the same kind of nutritive roots so well known 

 in Dahlia, Ficaria, etc. The plant is perennial and the germ- \ 



ination shows a few points of interest. A germinating plant- _^ 



let has been figured on Plate V, Fig. i, where we see the two m 



long-petioled cotyledons, the blades of which are ovate and 

 almost obtuse (Fig. 2). There is no distinct hypocotyl,* but 

 the primary root, which seems to commence immediately m 



under the cotyledons, is strongly developed and shows a cer- 

 tain ability of growth in thickness in this very early state. 

 The end of the root is sometimes bifurcated and terminates 

 suddenly in thin branches. The first leaf shows the general 

 features of the final ones, but has merely a smaller number of 

 divisions. A somewhat older state is shown in Fiij. %. where 



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