A 



B. 



<1% 

 1% 

 1 .5 % 

 2% 



n = 47 



n = 18 



22 



<1% 1% 1.5% 2% 2.5% 3% 

 Gradient 



n = 9 



Figure 8: Stream gradients at sample sites containing A) slimy sculpins, B) torrent 

 sculpins, C) shorthead sculpins and D) gradients for the total number of sites sampled. 



