210 MR. F. E. BEDDARD ON A 



which is quite continuous above with the lining of the genital 

 cloaca and below with the epithelium of the cirrus sac. The 

 epithelium appears to be quite the same throughout this whole 

 tract. It is clear, in fact, that both the narrow cirrus and 

 the flask-shaped pouch into which it opens form one structure 

 possibly invaginated from the genital cloaca, which is quite distinct 

 morphologically from the sperm-duct. The cirrus sac itself is 

 also divisible into two regions when it is occupied by the fully 

 retracted cirrus, as shown in text-figure 27. The almost spherical 

 distal part leads through a narrower neck to the cloaca genitalis. 

 The spherical bulb is covered externally by a thin layer of muscle- 

 fibres. The interior of the sac is filled with a tissue like that seen 

 in the cirrus sac of other tapeworms, for example, in Anoplotmnia, 

 where I have represented its principal characteristics*. 



There are many nuclei belonging apparently to delicate muscular 

 fibres and forming a padding tissue between the contained coils 

 of the male efferent system. Running from the wall of the 

 cirrus sac furthest from the pore are muscular slips which are 

 attached to the nearest end of the wide sac into which the cirrus 

 opens and are doubtless retractile in function. The narrower 

 neck-part of the cirrus sac is tubular and has a much thicker 

 coating of muscles, which are circular in direction. This layer is 

 one with the thin muscular coat of the bulb-region of the cirrus 

 sac. Between this thick muscular coating and the terminal part 

 of the male efferent apparatus which lies within it is seen a 

 delicate muscular layer with abundant interspersed nuclei like the 

 packing tissue of the bulb of the cirrus sac. The fibres, however, 

 have a definitely longitudinal direction, and would seem to be 

 part of the retracting apparatus. 



It has been mentioned that the cirrus sac projects into the 

 cloaca genitalis. The surface which thus projects is covered on 

 that part of it which faces the external pore by close chevaux 

 de /rise of minute conical spines, which are not implanted hooks 

 such as often occur upon the cirrvis in tapeworms, but rather 

 appear to be outgrowths of the cuticular covering. They suggest 

 to some extent a comparison with a group of fine setse described 

 by Fuhrmannf in the two species Anomotcenia penicillata and 

 A. isacantha in an apparently similar situation. 



The vas deferens immediately on leaving the cirrus sac forms a 

 dense coil filling up a good deal of space. The tube is here wider 

 than the sperm-duct within the cirrus, and the nuclei of the cells 

 which constitute its walls are very obvious ; the cells themselves 

 are rather clear. I could find no vesicula seminalis along the 

 course of the sperm-duct. 



The uterus of this worm, together with its contents — the ripe 

 or ripening embryos — shows certain definite peculiarities. At its 

 first appearance the uterus is sac-like and narrow, lying in front 

 of the ovaries and testes but still not very near to the anterior 



» P. Z. S. 1911, p. 1015, text-fig. 215. 



t Centralbl. f. Bakt. u. Parasit. xlv. 1908, p. 516. 



