928 MR. R. I. POCOCK ON THE FEET 
use of the fore feet, partly in the interests of brevity, partly because 
they supply the best characters, and partly because the features 
presented by the vibrissee and rhinarium call for verification on 
fresh material, which is unavailable at the present time :— 
a. Pad of pollex set low down close to the posterior lateral angle 
of the plantar pad; pads of third and fourth digits basally 
united; edge of web between median and lateral digits naked ; 
area between digital and plantar pads and between carpal and 
plantar pads scantily covered with short hair ............ Speothos venaticus. 
a’. Pad of pollex, when present, high above plantar pad ; pads of 
third and fourth digits separated; edge of webs joing the 
median with the lateral digits hairy ; area between digital pads 
and plantar pad and between the latter and the carpal pad 
mostly thickiy hairy. 
b. Edge of web joining third and fourth digits naked, forming a 
definite hairless band; a long fringe rising behind this and 
extending round the periphery of the foot behind the digital 
pads. 
ec. Pollex suppressed, feet long, carpal pad high above plantar 
pad ane cin Sa < bbe eal. Seek ames Lycaon pictus. 
ce’. Pollex retained; feet shorter, carpal pad moderately high 
above plantar Pads. .<s.cn2.-c-con-nteerectbecercsesersmssensass CULO POPU GMS: 
b’. Edge of web between third and fourth digits hairy. 
d. Glandular depression between pads undivided ; integument 
proximal to the pads not specially thickened. 
e. Third and fourth digits exceedingly widely separable, the 
edge of their web when stretched forming a straight line 
about twice the length of the pads of either digit; plantar 
pad long and narrow .............00600e00:c01eeeee Pseudalopex gracilis. 
e’. Third and fourth digits much less widely separable, the 
edge of the web between them emarginate when stretched 
and less than the length of the pad of either digit. 
f. Plantar pad large, its width exceeding the length between 
its median lobe and the margin of the web between the 
shorter third and fourth digits; claws short. 
g. Carpal pad very small, third and fourth digits more 
widely separated, the web joining them, when ex- 
tended, as wide as that between them and the 
Nateralidigiisy s.ieecee selec cstaecs Cerdocyon microtis. 
g’. Carpal pad large, third and fourth digits more closely 
united; the connecting web narrower than that 
between them and the lateral digits... Canis anthus, mesomelas*. 
f’. Plantar pad small, its width considerably less than the 
distance between its median lobe and the edge of 
the narrow web between the long third and fourth 
digits; claws long vecereccesscsseeeese Vulpes vulpes and 
bengalensis ; Otocyon megalotis. 
d’. Glandular depression between digital and plantar pads 
divided by a ridge of thick skin stretching forwards from 
the median lobe of the plantar pad; integument surround- 
ing proximal portion of digital pads much swollen and 
cushion-like; feet otherwise almost as under e’ ......... Alopex lagopus. 
The above-given analysis of the characters of the feet of the 
Canidee emphasises the distinctness. of Speothos from the rest of 
the genera, and does not afford support to the affiliation of Speothos 
* The feet of domestic breeds of dogs come under this heading. The feet of 
some breeds indeed conform very closely to the type seen in Canis anthus (see 
P. ZS. 1914, pp. 478-484). 
