1901.] ANATOMY or COGIA BEEVICEPS. 123 



No doubt this matter is dealt with by the French authors ; but 

 in view of the condition of the " stomach " in other Oetacea, that 

 of the Cachalot is rather puzzling. 



It appears, then, that in the family Physeteridae the apparatus is 

 simpler than in the Delphinidae, and agrees with the Platanistidae in 

 having only three chambers, viz., a paunch, a cardiac chamber, and 

 a pyloric chamber. 



V. The Penis. 



The body- wall between the anus and the aperture of the penial 

 sheath had been cut about, so that the position of these apertures 

 and their distance apart, with regard to the body-length, could not 

 be ascertained. 



In the arrangement of the muscles at the base of the organ, and 

 some other matters, Oogia differs from the accounts given for the 

 Dolphin and the Eight Whale'(by Professors Turner and Struthers 

 respectively), so that it is worth while to put on record the arrange- 

 ments in the present whale. The total length of the apparatus, in 

 the fresh condition, was 28 inches, measured from the base of the 

 " accelerator urinjB " muscles to the end of the penial sheath, but 

 more detailed measurements of the preserved material show that 

 a certain amount of shrinkage had occurrred. 



In dealing with the penis, it will be convenient to distinguish 

 three regions : — 



(1) A basal portion consisting of the crura penis, surrounded by 



the muscles, which in Oogia are 4 in number, a pair of 

 medial acceleratores urince and a pair of lateral erectores penis. 

 This region measures, in the preserved specimen, 7 inches in 

 length and about 5 inches at the broadest. 



(2) The middle region or " body of the penis," rather more than 



6 inches in length, formed almost wholly by the single 



corpus cavernosum. 



And (3) the distal region of 12 inches, extending from the 



insertion of the sheath or prepuce into the penis, up to the 



external pore. The terminal region of the penis, enclosed 



within the sheath, is 8| inches in length : this region may 



be termed the ' glans,' though it contains a continuation of 



the corpus cavernosum. 



We will consider the middle region or body of the penis first. 



It consists of a single corpus cavernosum, having the usual struc- 



ure, with a thick tunic of fibrous tissue, and enveloped in a looser 



connective tissue carrying blood-vessels and nerves (PL X. fig. 16), 



The body is not quite cylindrical, but is slightly higher than broad 



(1| inch by 11 inch, in the measured specimen) ; it was not straight, 



but somewhat undulating ^. 



The upper surface is convex. There is no dorsal furrow, such 

 as is described for Balcenoptera, and the plexus of blood-vessels 



^ The only figure of transverse sections of a Cetacean penis that I have seen 

 is that given by Murie (4) for Balmnoptera, where it is broader than high. 



