916 MR. F. E. BEDDARD ON THE ANATOMY [Dec. 15,.. 



process arises which runs forward as far as the angle of the lower 

 jaw, which has almost the appearance of a duct, but which is, of 

 course, no such thing. The glands are white and appear to be 

 chiefly composed of fat. 



In addition to these structures there is on each side of the body 

 another " gland " of the same general appearance, but smaller, 

 which I refer to the same category. These are flat circular bodies 

 2*5-3 mm. in diameter. These lie on either side well behind the 

 head and just behind the scapula, covered over by the fascia which 

 is continued into the depressor mandibulfe, and have a special 

 relation to the thickened lateral edge of that muscle, as is described 

 elsewhere *. Their position is considerably more posterior than 

 the thymus glands of Rana and than the additional glands of 

 Hemisus already described. It is well known f that among the 

 Amphibia there are in the adult various remains of bodies derived 

 from cells belonging to the branchial system of the tadpole, which 

 have been termed " postbranchial bodies," " Epithelkorperchen," 

 &c. I take it that in Hemisus these various structures which 

 have just been described are also to be referred to the same 

 category. They are, however, unquestionably much larger rela- 

 tively as well as a.ctually than in at least some other Amphibia 

 Salientia. 



§ Posterior Lymph-Hearts and Associated Sacs. 



The enormous size, actual as well as relative, of the posterior 

 lymph-hearts of Brevicej)sX is at present an unique fact in the 

 anatomy of the Batrachia Salientia. On grounds of affinity the 

 existence of equally or nearly as large posterior lymph-hearts 

 might have been expected in Khinodervna darwini ; but a careful 

 search convinced me that Rhinoderma § was unlike Breviceps: 

 in this important and remarkable peculiarity. In Hemisus,. 

 however, I find an equivalent of this structure, which is very 

 different in its character, though i-etaining certain features which 

 lead to the inference that we have in this genus a modified 

 and degenerate homologue of the enormous posterior lymph-heart, 

 of Breviceps verrucosus. 



As is the case with Brevicejos, though not to so great an extent, 

 as in that Frog, the thigh of Hemisus is fairly enclosed within 

 the contour of the body. The body does not, however, extend 

 beyond the tip of the coccyx as it does in Breviceps. 



There is thus in Honistos, as in Breviceps, a space left dorsally 

 on each side of the posterior region of the vertebral column, 

 behind the oblique muscles, which is floored by the muscles of the- 

 proximal region of the thigh. This space is cut off from the 

 lymph-spaces covering the rest of the thigh by a transverse and 



* Cf. p. £94. 



t Maurer, " Schildru.se, Thymus uiid Kiemenreste der Ampliibien," Morph. Jahrb^ 

 xiu. 1888, p. 298. 



t Beddard, " Anatomy of Breviceps," P. Z. S. 1908, p. 33. 

 ^ § Id., " On Rhinoderma," P. Z. S. t. c. p. 678. 



