922 MR. F, E. BEBBARD ON THE ANATOMY [DeC. 15, 



large lobate fat-body already described and figured (text-figs. 182, 

 183, pp. 917, 919). At the same time it will be noted that the 

 *' chordte tendineae " of the lymph-sac of Eana would seem to fore- 

 shadow (or to be the remains of) this trabecular system. The 

 mass of trabeculaj and contained fat-mass is carried to an extreme 

 in the opposite direction in the case of Xenojnis, where (see 

 p. 924) a tough mass of connective-tissue fibres and intersjoersed 

 fat surrounds the lymph-hearts. 



It is said, and in a sense correctly, that there are no lymphatic 

 glands in the Frog. The absolute truth of this generalization 

 (which of course applies to other lower Vertebrates as w^ell as the 

 Frog) depends upon what is meant by the term " Ij-mphatic gland." 

 For if we legard a lymphatic gland as an enlai-gement on the 

 covirse of a lymphatic vessel the lumen of which is subdivided by 

 trabeculfe, then the structures described here in Hemisiis are at 

 least not very unlike lymph-glands. One cannot but think, in 

 view of the masses of fatty tissue with which they are partly 

 plugged, that they must play some important part in the function 

 of the lymphatic system ; and the existence of fatty masses in 

 Xenojms, described below *, strengthens this supposition in that 

 it shows that the structure is not unique. 



The lymph-heart itself is of considerable size, though not 

 so colossid as in Bt^eviceps. It measures fully 4 mm. in length 

 and is rather elongate in form. It is displayed in text-fig. 186, l.h., 

 from which it will be seen that the heart occupies quite a normal 

 position. It lies, however, quite definitely within the large 

 saccus iliacus which has just been described. In Bcma escidenta 

 the lymph-heart of the same pair is said by Gauppt to lie in 

 a special sac of its own. This is certainly not the case with 

 Hemisus. The posterior lymph-heart lies at the inner angle of 

 the saccus iliacus. The heart is in contact with the vastus 

 externus, upon which it lies ventrally ; with the ilio-coccygeal 

 muscle to the inside. On the opposite side its wall lies freely 

 within the lymph-sac already mentioned and described at length. 

 It is in front of the pyriformis muscle. It is not very closely 

 related to the giutfeus muscle. The septum bounding the saccus 

 iliacus posteriorly is attached along the vastus externus and ends 

 on the wall of the heart, or at least is firmly attached to the 

 posterior wall of the heart at this point. Just in the angle 

 where the septum in question and the heart meet is a smallish 

 circular ostium (text-fig. 186, 0.), which I take to be a communi- 

 cation between the saccus iliacus and the heart. A small blood- 

 vessel, as is shown in the figure {n), runs along the outer side of the 

 heart forwards, dipping down to the gluteal "muscle; it is possible 

 that this is the vein into which the heart opens. The heart is 

 rather pear-shaped and lies with its long axis parallel to the long 

 axis of the body. The narrow end is anterior. The heart is easily 

 to be distinguished from the posterior septum of the saccus iliacus 



* See p. 924. + See his edition of Ecker's ' Fi-oa:.' 



