A NEW REPTILIAN TREMATODE, 935 



end, where it is surrounded by large deeply staining oells ; just 

 in front of the opening the vesicle is much folded, giving it 

 the appearance of having been " puckered in " before the narrow 

 opening. In front of this pleating it widens out as a large tube 

 running dorsal to and parallel with the ascending limb of the 

 uterus ; in front of the testes and behind the ovary it forks, each 

 limb reaching far forward beyond the ventral sucker to about 

 midway between the ventral sucker and pharynx. 



The testes are oval bodies more or less lobed on their inner 

 margins, usually lying obliquely with the right testis in front of 

 the left ; the usual situation is for the right to be about 0'15 mm. 

 in front of the left, but they may be almost on a level, and in 

 two cases the left testis was in front of the right. Curiously 

 enough, these variations occurred in the small immature worm 

 and in the largest specimen. It is just possible that we may 

 have to do here with a different species. In a specimen 5'2 ram. 

 in length the testes measured, right 0*90 mm. x 0-44 mm., left 

 0"84 mm. x 0*44 mm,, the greater axis lying longitudinally. On 

 the inner margin there are two or three lobes. The cii-rus-sa.c is 

 long and reaches almost to the posterior leA^el of the ventral 

 sucker. Sometimes it reaches the extreme posterior end, and in 

 two cases (again the immature worm and the largest) it was 

 much shorter and only reached a short way behind the antei'ior 

 margin of the ventral sucker ; in this respect being much nearer 

 L. validus IS^icoll. The vesicula seminalis is much convoluted 

 and occupies the posterior third of the cirrus-sac, the ductus 

 ejaculatorius being surrounded by an enormous mass of prostatic 

 cells and ending in a short protrusible cirrus often to be seen 

 exserted from the genital pore. The male opening is on the 

 right side of the genital pore, which is situated on a level with 

 the centre of the pharynx and midway between it and the lateral 

 margin of the body-wall. The ovary is almost round, and lies 

 dorsally on the right side of the body immediately in front of the 

 right testis. It overlaps the ventral sucker for about a third of 

 its diameter ; its position varies however, and in some cases it 

 may be rather more than this and in others not so much. The 

 portion of the ovary that overlaps the cirrus-pouch is on its right 

 side. The oviduct receives a large shell-gland, gives off a Laurer's 

 canal, and receives the vitellai-ian duct, then runs down as the 

 uterus in many coils to nearly the posterior end of the body, 

 when it doubles back as a much broader tube, runs forward 

 gradually straightening and narrowing and passes the ventral 

 sucker dorsally. In the course of the descending portion behind 

 the right testis is a receptaculum seminis uterinum. When the 

 uterus reaches the level of the anterior ends of the excretory 

 vesicle its walls form a vagina with a thick cuticular lining, and 

 surrounded by a layer of deeply staining large cells which 

 accompany it for the whole of its course until it opens on the 

 left side of the genital pore, The eggs are very numerous and 

 of a deep brown colour when old, the young eggs being much paler. 



