158 DR. F. E. BEDDARD ON 



medullary parenchyma is more dense. In the fully mature 

 segments towards the end of the body both the cortical and 

 the medvillary parenchyma are equally lax in structure. This 

 renders it very difficult to follow the excretory tubes the calibre 

 of which is not greater than that of the spaces between the fibres 

 of the parenchyma. 



This species also contrasts with some others (as is shown in 

 text-figs. 33 & 34) in the large and very conspicuous calcareous 

 corpuscles. 



The testes are numerous and show the usual arrangement met 

 with in this genus. In ripe segments I counted as many as ten 

 testes in one transverse row, five on either side of the median 

 uterus. It appeared to me that in this species the entire genera- 

 tive system was ripe at the same time. That is to say, the testes 

 were mature in segments in which the uterus was filled with 

 eggs ; and, on the contrary, more anterioi-l}^, where the uterus 

 was only represented by a slender median cord, in which a lumen 

 was haiTlly discernible, the testes were also immatvire. They 

 are at first represented by patches of nuclei in the medullary 

 parenchyma. Later they are in the form of sharply marked 

 cavities, loosely packed with the testicular cells, and when fully 

 mature the masses of spermatozoa nearly fill the cavities. The 

 sperin-duct forms the usual coil close to its opening into the 

 cirrvis-sac. This coil appeared to me to be denser than in 

 some other species, and the lumen of the sperm-duct not to 

 be quite so wide as is often the case. The coil of the sperm- 

 dvict and the cirrus-sac together occupy from one-third to 

 one-half of the diameter of a proglottid, and lie at about its 

 middle, sometimes rather anterior, at times rather posterior, to 

 the middle line. 



The cirrus sac is, as a rule, almost spherical in shape in the 

 mature proglottids. Its walls are thin. The cirrus itself lies 

 coiled up within it, and not much room is left between the cirrus 

 and the walls of the sac in which it lies. Everywhere the cirrus 

 is surrounded by a layer of glandular cells like those to which I 

 have referred in Ophidotcenia naice *. When the cirrus-sac is 

 examined in an entire proglottid mounted in glycerine, the 

 anterior and outside region of the sac is seen to be occupied by 

 the distal part of the cirrus, which is of much greater calibre 

 than the rest and lies almost straight. The rest of the sac is 

 occupied by the close coils of the narrower region of the cirrus. 

 In such preparations the cirrus is of a golden yellow colour, and 

 thus stands out conspicuously from the rest of the tissues of the 

 Avorm, I have never observed the cirrus in a state of protru- 

 sion ; but, on the other hand, I have seen the whole sac itself 

 partially protruded in a way commented upon by Schwarz f in 

 other species of this genus. I found no spines upon the cirrus 

 anywhere. The vagina runs at first straight and parallel with 



* P. Z.S. 1913, p. 29. 



t " Ichthyotsenien d. Reptilien," Inaug.-Diss. Basel, 1908. 



