REPTILIAN TAPEWORMS. 259 



these ventral openings is mentioned ; thus, e. g., he writes of 

 Proteocephalus singidaris, sp. n. : " Uterus with 20-25 lateral 

 pouches on either side and with 2-4 ventral pores." The same 

 statement (save for the actual number of the pores) is made of one 

 species of Ophiotamia *, a new genus described by the author. 



This comprises all that is written in the preliminary note 

 concerning the uterine pores. There is enough, however, as I 

 believe, to prove that Mr. La Rue and myself have observed 

 identical structures; but it is, on the other hand, conceivable that 

 the conditions characteristic of the various species described by 

 La Rue are similar to those which I have described above in 

 Solenota^nia, and that the "ventral openings" of the uterus may 

 be merely the commencement of ruptures which end in fusing 

 together and forming a ventral groove. This view would account 

 for the variable num]:)er of the openings in a given species, of 

 which I have quoted an instance above. 



The testes of this species are very numei'ous, and really extend 

 quite across the body except for the middle regon occupied by 

 the uterus, aiad this is relatively quite narrow. Posteriorly the 

 testes reach the ovary, and are found at the side of that gonad 

 which is not greatly extended laterally. The most posterior 

 region of the proglottid is, however, fi-ee of testes ; not however 

 the most anterior, for they extend quite to the anterior limit of 

 the proglottid. In younger proglottids, where the uterus is 

 simply represented by a narrow median cord, the testes are more 

 obviously disposed in two lateral bands, leaving a considerable 

 gap in the region of the uterus, which is to be occupied by the 

 latter when fully mature and provided with its diverticida. 

 In mature proglottids I have seen quite ten testes on either 

 side of the uterus ; but they are not arranged in regular rows. 

 The coil of the vas deferens before it enters the cirinis-sa,c is 

 denser than I have seen it in some allied forms. The coil 

 extends quite to the uterus, and the lumen of the tube is wide. 

 The cirrus-sac (text-fig. 53) seems hardly to possess a definite wall 

 of its own. The wall of the cavity in which the coiled cirrus lies is 

 practically a layer of the network of the medullary parenchyma. 

 This region of the cirrus-sac is hardly wider than that situated 

 nearer to the external pore. Further in the latter direction the 

 bands of longitudinal muscle-fibres come to lie actually on the wall 

 of the cirrus where, however, they are obviously merely adven- 

 titious and undergo no modification, such as increase in thickness, 

 to assist in the protrusion of the cirrus. The cirrus-sac is not so 

 markedly flask-shaped as it is in many other Cestodes ; but there 

 is a narrower neck region towards the external pore. This region 

 has marked but not very thick muscular walls, in which the 

 fibres trace a more or less circular direction, thus forming a 



* I do not know whether this genus is exactly coextensive with my {jenus Opliido- 

 tcenia. The preliminary definition given of Ophiotmnia is not sufficient. The 

 accidental similarity of names is a coincidence which might well have been expected. 



