STATE OF ILLINOIS DEPARTMENT OF REGISTRATION AND EDUCATION 



BOARD OF NATURAL RESOURCES AND CONSERVATION 

 John C. Watson, Chairman; Thomas Park, Ph.D., Biology; L. L. Sloss, Ph.D., Geology; Roger Adams, Ph.D 

 D.Sc, Chemistry; Robert II. Anderson, B.S.C.E., Engineering; Charles E. Olmsted. Ph.D., Forestry; W. L 

 Everitt, E.E., Ph.D., Representing the President of the University of Illinois; Delyte \V. Morris. Ph.D., Preside?, 

 oj Southern Illinois University. 



NATURAL HISTORY SURVEY DIVISION, Urbana, Illinois 



SCIENTIFIC AND TECHNICAL STAFF 



Harlow B. Mills, Ph.D., Chief 



Herbert H. Ross, Ph.D., Assistant Chief 



Robert O. Watson, B.S., Assistant to the Chief 



Section of Economic Entomology 



William II. Luckmann, Ph.D.. Entot Head 



Willis N. Bruce, Ph.D., Entomologist 

 Ronald 11. Meyer, Ph.D., Associate Entomologist 

 Iames E. Appleby, Ph.D., Associate Entomologist 

 Robert D. Pausch, Ph.D., Assistant Entomologist 

 Ralph E. Sechriest, Ph.D., Assistant Entomologist 

 Delmar Broersma, Ph.D., Assistant Entomologist 

 Joseph V. Maddox, Ph.D.. Assistant Entomologist 

 Edward J. Armbrust, Ph.D.. Assistant Entomologist 

 Dannel McCollum, B.A., Technical Assistant 

 Gregory P. Marsh. B.S., Technical Assistant 

 Rami Banerjee, B.A.. Technical Assistant 

 Alan C. York. B.S.. Technical Assistant 

 Sue E. Watkins, Junior Scientific Assistant 

 H. B. Petty, Ph.D., Entomologist in Extension* 

 Stevenson Moore, III. Ph.D.. Entomologist in Exten- 

 sion* 

 Roscoe Randell, M.S., Technical Assistant in Exten- 

 sion* 

 Clarence E. White, B.S., Technical Assistant in Exten- 

 sion* 

 Stanley Rachesky, M.S., Technical Assistant in 



Extension* 

 D. E. Kuhlman, B.S.. Technical Assistant in Extension* 

 Amal C. Banerjee, Ph.D., Research Associate* 

 Jean G. Wilson, B.A., Research Associate* 

 Ayten Hatiboglu, B.S.. Research Assistant* 

 Keun S. Park, M.S., Research Assistant* 

 Keturah Reinbold, M.S., Research Assistant* 



Section of Faunistic Surveys and 

 Insect Identification 



H. H. Ross, Ph.D., Assistant Chief and Head 



Milton W. Sanderson, Ph.D., Taxonomist 



Lewis J. Stannard, Jr., Ph.D., Taxonomist 



Philip W. Smith, Ph.D., Taxonomist 



Leonora K. Gloyd, M.S., Associate Taxonomist 



Wallace E. LaBerge, Ph.D., Associate Taxonomist 



Robert T. Allen, M.S., Technical Assistant 



Bernice Sweeney, Technical Assistant 



Bess White, A.B., Technical Assistant 



John D. Unzicker. Ph.D.. Research Assistant* 



Toshio Yamamoto, M.S., Research Assistant* 



Section of Aquatic Biology 



George W. Bennett, Ph.D., Aquatic Biologist and Head 



William C. Starrett, Ph.D., Aquatic Biologist 



R. W. Larimore, Ph.D., Aquatic Biologist 



David H. Buck, Ph.D., Associate Aquatic Biologist 



Robert C. Hiltibran, Ph.D., Associate Biochemist 



Donald F. Hansen, Ph.D., Associate Aquatic Biologist 



William F. Childers. Ph.D., Assistant Aquatic Biologist 



David L. Thomas, B.S., Research Assistant 



Maryfran Martin, Technical Assistant 



Claude Russell Rose, Field Assistant 



Charles F. Thoits, III, B.A., Research Associate* 



Section of Applied Botany and Plant Pathology 



J. Cedric Carter, Ph.D., Plant Pathologist and Head 



I. L. Forsberg, Ph.D., Plant Pathologist 



Robert A. Evers, Ph.D., Botanist 



Robert Dan Neely, Ph.D., Plant Pathologist 



E. B. Himelick, Ph.D., Plant Pathologist 



Walter Hartstirn, Ph.D., Assistant Plant Pathologi 



D. F. Schoeneweiss. Ph.D., Assistant Plant Pathologi 



Bruce L. Johnson, B.S., Technical Assistant 



David Russell Vance, Technical Assistant 



Robert J. Slattery, B.S., Technical Assistant 



Robert S. Perry, B.S.. Technical Assistant 



Betty S. Nelson, Technical Assistant 



Sylvia P. Brockstein, M.S., Technical Assistant 



Section of Wildlife Research 



Glen C. Sanderson, Ph.D., Wildlife Specialist and Head 

 F. C. Bellrose, B.S., Wildlife Specialist 

 H. C. Hanson, Ph.D., Wildlife Specialist 

 Richard R. Graber, Ph.D., Wildlife Specialist 

 Ronald F. Labisky, M.S., Associate Wildlife Specialist 

 William R. Edwards, M.S., Associate Wildlife Speciali 

 William W. Cochran, Jr., Assistant Wildlife Speciali 

 Rorert E. Greenberg, M.S., Research Assistant 

 Jeffrey C Hanson, M.S., Research Assistant 

 Helen C. Schultz, B.S., Technical Assistant 

 Lucinda Janis, Technical Assistant 

 Robert D. Crompton, Field Assistant 

 William J. Francis, Ph.D., Research Associate* 

 Jack A. Ellis, M.S., Research Associate* 

 James A. Bailey, Ph.D., Research Associate* 

 Gerald G. Montgomery. M.S., Research Associate 

 William L. Anderson, M.A., Research Associate* 

 George B. Joselyn. M.S., Research Associate* 

 Gerald L. Storm, M.S., Field Ecologist* 

 Ronald L. Westemeier, B.S., Research Associate* 

 Stanley L. Etter, M.S., Research Assistant* 

 Robert E. Hawkins, B.S., Research Assistant* 

 Larry D. Martoglio, M.S.. Research Assistant* 

 Keith P. Dauphin, Project Assistant* 



Section of Publications and 

 Public Relations 



Owen F. Glissendorf, M.S., Technical Editor and He, 

 Robert M. Zewadski, M.S., Associate Technical Edit 

 Nancy K. Ginzel, B.S., Assistant Technical Editor 

 Wilmer D. Zehr, Technical Photographer 

 William L. Taylor, Technical Assistant 



Technical Library 



Doris F. Dodds, B.A., M.S.L.S., Technical Librarian 

 Patricia F. Stenstrom, B.A., M.S.L.S., Assistant Tech- 

 nical Librarian 



CONSULTANTS: Herpetology. Hobart M. Smith, Ph.D., Professor of Zoology, University of Illinois; ParasitolOC 

 Norman D. Levine, Ph.D., Professor of Veterinary Parasitology and Veterinary Research, University of Illinoi 

 Wildlife Research, Willard D. Klimstra, Ph.D., Professor of Zoology and Director of Cooperative Wildlife R 

 search, Southern Illinois University; Statistics, Horace W. Norton, Ph.D., Professor of Statistical Design ai 

 Analysis, University oj Illinois. 



♦Employed on cooperative projects with one of several agencies: University of Illinois, Illinois Agricultural Exte 

 sion Service, Illinois Department of Conservation, National Science Foundation, United States Department of Agi 

 culture, United States Fish and Wildlife Service, United States Public Health Service, and others. 



This paper is printed by authority of the State of Illinois, IRS Ch. Ill . Par. 5S.12. It is a contribution from t. 

 Section of Applied Botany and Plant Pathology of the Illinois Natural History Survey. 



Illinois Natural History Survey Circular 52, September, 1966 



(31847— 10M— 9-66)30 



