NEW TAPEWORMS FROM THE HTRAX. 583 



in Thysanotcenia gambiana. The point of difference, however, is 

 not a large one, since in other species of the present genus there 

 is no difi'erentiation of the testes into two masses, but they form 

 a continuous mass reaching from side to side of the proglottid. 

 In the case of the group of testes on the pore side, they lie to the 

 pore side of the ovary and vitelline gland. These testes were 

 very conspicuously pear-shaped, the duct naturally arising from 

 the pointed end. In the case of the testes on the pore side, at 

 any rate, the narrower end faced forwards in the segment. In the 

 case of each group the testes are closely packed together. They 

 are restricted to the medullary part of the parenchyma. The 

 efferent sperm-ducts from the testes which lie away from the 

 pore side form a widish trunk which breaks off and forms a slight 

 rete mirabile not so marked as in the species next to be described. 

 This opens evidently by one thick tube into the large vesicula 

 seminalis lying close to the ovary and to the receptaculum seminis. 



The cirrus sac is not conspicuously large, and it lies close to 

 the lateral margin of each proglottid and thus well within the 

 cortical parenchyma. Its position is oblique, the end of the 

 sac being anterior to its aperture on to the exterior, and is 

 thus better shown in horizontal than transverse sections of the 

 proglottids. The general outline of the cirrus sac, when displayed 

 in such sections, is, like that of many other species (e. g. 

 Otiditcenia eupodotidis *), flask-shaped, there being a narrower 

 proximal neck and a wider distal region. It lies parallel to the 

 vagina and very close to it, but in a different plane, so that a 

 single horizontal section does not display both tubes lying side 

 by side. Such a section, however, does show the extreme 

 terminal part including the external orifice of both cirrus sac and 

 vagina ; and it will be seen that they lie accurately side by side, 

 the male pore being, of course, anterior to the female pore. Such 

 a section also shows very plainly that there is no trace of a 

 cloaca genitalis ; the two pores are upon the surface of the body 

 upon a slightly raised papilla of not very great dimensions. The 

 papilla which bears the two orifices is clearly of less importance 

 than that which charactei'ises the two worms for which I have 

 recently formed the genus Thysanotcenia f. On a re -examination 

 of my sections of the species Thysanotcenia lemuris I find that 

 in that species, as well as in that which forms the subject of the 

 present communication, there is no sinus genitalis, but that both 

 cirrus sac and vagina open side by side on the summit of the 

 papilla. The case is otherwise with Thysanotcenia gcombiana, 

 where there is a distinct though not very large sinus or cloaca 

 genitalis. There are also corresponding differences in the vagina 

 and cirrus sac of the latter, which I shall deal with after describing 

 the ari'angements found in the Anoplocephalid from the Hyrax. 



The cirrus sac has a thickish muscular coat which does not 

 become at all markedly thicker in the "neck" region. In 



* P. Z. S. 1912, p. 206, text-fig. 27. 

 t P. Z. S. 1911, p. 1002. 



