EPIPHYLLUM. 



Hermann (Par. Botavus Prodr. Add. 2 



he listed the name Epiphyllum americanum. 

 in 18 12 when he estabHshed the genus. 



187 



1689) first used the name Epiphyllum when 

 Haworth credited the name also to Hermann 



Key to Species. 



A. Perianth-tube 7 to 9 times as long as the limb i. £. phyllanthus 



AA. Perianth-tube 1 1 2 to 3 times as long as the limb. 

 B. Ultimate joints acuminate. 



Flowers 25 to 30 cm. long 2. E. oxypetalum 



Flowers 10 to 15 cm. long. 



Margins of joints crenate ; 3. E. pumilum 



Margins of joints undulate 4. E. caudatum 



BB. Ultimate joints acute, obtuse or rounded. 

 C. Joints deeply lobed. 



Joints 2 to 7 cm. broad. 



Lobes of joints spreading; outer perianth-segments lemon- yellow 5. -E. darrahii 



Lobes of joints pointing forward; outer perianth-segments reddish yellow. . 6. E. anguliger 



Joints very large, up to 25 cm. broad 7. E. grandilobum 



CC. Joints crenate or nearly entire. 



D. Joints deeply crenate, thick; perianth-tube bearing foliaceous scales &. E. crenatum 



DD. Joints low-crenate to nearly entire; perianth-tube without foliaceous scales. 

 E. Sinui of the joint-margins very narrow; flowers up to 20 cm. broad; 



stamens yellow 9. E. mac opterum 



EE. Sinui of the joint-margins open; flowers 15 cm. wide or less; stamens 

 not yellow. 



Ovary and fruit bearing linear scales 10. E. lepidocarpum 



Ovary and fruit without linear scales. 



Flowers 10 to 13 cm. long 11. E. pittieri 



Flowers 15 to 28 cm. long. 



Flowers about 28 cm. long; style orange 12. E. guatemalense 



Flowers 15 to 25 cm. long; style white or pink. 



Joints very stiff 13. E. stridum 



Joints flexible or moderately stiff. 



Joints very large, up to i meter long and 12 cm. wide 14. £. stenopetalum 



Joints smaller, rarely ever 7 cm. wide. 



Joints shallowly crenate or subdentate; species of Costa 



Rica 15. E. carlagense 



Joints deeply crenate; species of Tobago, Trinidad, and 



Venezuela 16. E. hookeri 



1. Epiphyllum phyllanthus (Linnaeus) Haworth, Syn. PL Succ. 197. 1812. 



Cactus phyllanthus Linnaeus, Sp. PI. 469. 1753. 



Opuntia phyllanthus Miller, Gard. Diet. ed. 8. No. 9. 1768. 



Cereus phyllanthus De Candolle, Prodr. 3: 469. 1828. 



Phyllocactus phyllanthus Link, Handb. Erkenn. Gewachse 2:11. 1831. 



Rhipsalis macrocarpa Miquel, Bull. Sci. Phys. Nat. Neerl. 1838: 49. 1838 (in most part). 



Rhipsalis phyllanthus Schumann in Martius, Fl. Bras. 4": 298. 1890 (in part). 



Hariola macrocarpa Kuntze, Rev. Gen. PI. i: 263. 1891. 



Phyllocactus phyllanthus paraguayensis Weber, Diet. Hort. Bois 957. 1898. 



Phyllocactus phyllanthus boliviensis Weber, Diet. Hort. Bois 957. 1898. 



Phyllocactus phyllanthus columbiensis Weber, Diet. Hort. Bois 957. 1898. 



Epiphyllum gaillardae Britton and Rose, Contr. U. S. Nat. Herb. 16: 240. 1913. 



Phyllocactus gaillardae Vaupel, Monatsschr. Kakteenk. 23: 87. 1913. 



Elongated and much branched; main branches narrow, terete or 3 or 4-angled, woody; termi- 

 nal joints elongated, terete or 3-angled below, usually flat or thin, rarely 3-winged, bright green with 

 a purple margin, sometimes 7 cm. broad, obtuse, the margin coarsely serrate, the teeth obtuse; 

 flower slender, 25 to 30 cm. long, the slender tube very much longer than the limb, green, the limb 

 greenish or white, its segments narrow, 2 to 2.5 cm. long; scales on flower-tube few, minute, spreading; 

 style long, slender, pinkish (Schumann says white) ; filaments short ; stigma-lobes short, white ; fruit 

 oblong, 7 to 9 cm. long, somewhat 8-ribbed, bright red; pulp white; seeds large, black, numerous. 



According to De Candolle, the flowers are nocturnal and odoriferous. 

 Type locality: Brazil. 



Distribution: Panama to British Guiana, Bolivia, Peru, and Brazil. Recorded from 

 Paraguay. 



