105 



last suggests failure in pollination, though there was nothing in 

 the outward appearance of the fruits to suggest so few nutlets 

 as 13 and 5. The proportion of undeveloped carpels to developed 

 was greater in the pyriform than in the napiform and globose, 

 but the total original number appeared to be fewer. Of this 

 last point I am not sure, as I neglected to examine flowering 

 specimens. It is worth noting that the same thing happens in 

 the elongated fruits of R. gymnocarpa, which commonly mature 

 only I or 2 carpels as compared with 4-8 in the globose fruits. 



The napiform and globose fruits thus mature the greatest 

 number of carpels. But that this does not determine the final 

 form is clear in the case of napiform fruits at least, from the 

 fact that their peculiar flat shape is quite easily recognized in 

 bud or flower. 



There is no concomitant relation between fruit-form and 

 prickle-form. Thus napiform fruits are accompanied by lanceo- 

 late (Fig. 7) broadly triangular (Fig. 9) and recurved (Fig. 8) 

 prickles; and pyriform fruits by either lanceolate or triangular 

 prickles. The flowers of napiform plants vary from 4.5-8 cm. 

 in width, and the stipules are either broad (Fig. 14) or medium 

 to narrow (Fig. 8). There is some variation in the glands of 

 sepals and peduncles, but if this has any significance I did not 

 detect it. 



Lastly these fruit-forms are not always so pure as in the Elgin 

 plants. Fig. i shows a common phenomenon, — the best- 

 developed fruits are mostly pyriform, the others subglobose. 

 (The large fruit at the right in this figure that appears to have 

 a round base, really has an acute base.) Necked and neckless 

 forms may occur on the same plant, as may napiform and globose; 

 while globose forms readily vary into ovoid or deltoid. While, 

 then, my observations result negatively, it is quite possible that 

 a wider survey may show greater significance in these forms 

 than I have been able to detect. The most important and 

 apparently the most widely distributed is the pyriform, which 

 shows a tendency to mature fewer nutlets than the globose or 

 napiform. 



Vancouver, B. C. 



