140 



(c) Similar to the above, but with less of the purplish tint 

 (more on spurs), and the whole flower (especially laminae) is 

 very pale yellow. Others show more of the purplish and brighter 

 yellow, intermediate between a and c. 



The factorial analysis of these forms cannot yet be clearly 

 made. A. caerulea in the western part of its range is not blue but 

 white, but there is no evidence that the plants we used carried a 

 recessive white. We can however postulate that yellow plastids 

 (chrysantha) are allelomorphic to their absence (caertdea), and 

 abundant anthocyanin (caerulea) to little or none (chrysantha). 

 It we call the factors respectively P, p, A, a, the Fi hybrid will 

 have the formula PpAa, and will combine blue with yellow, as 

 it actually does, with non-acidity also dominant over the acid 

 condition of chrysantha. In the F2 9 out of 16 should look like 

 theFi; three should resemble caerulea, three chrysantha, and one 

 might be expected to be white, feebly or not tinted with antho- 

 cyanin. Evidently other factors are involved, for as a matter 

 of fact the pallid (supposedly double recessive) flowers are 

 numerous. 



Genuine A. caerulea produces some hitherto unrecorded 

 variations. Mr. D. M. Andrews has at Boulder, Colorado, a 

 large stand of very fine and typical caerulea, the seed having 

 been obtained from the Blanchard ranch in Boulder Canon. 

 The strain originated in the nearby mountains, and is in general 

 extremely uniform. But as Mr. Andrews pointed out to me, 

 there are a few plants abruptly and conspicuously varying from 

 the type: 



1. Laminae of petals blue like the sepals, elongate, narrow 

 (e.g., 40 mm. long and 8 wide); spurs normal, varying to small 

 and more or less aborted. A few plants. This is more or less 

 intermediate between the typical form and variety daileyae, 

 but distinct from both. 



2. Flowers very pale, light yellowish or greenish in bud, 

 eventually delicately tinted with, purplish. Sepals and petals 

 9 to 10, the sepals reflected at maturity, placed just below the 

 outwardly-turned spurs; laminae remaining erect, lanceolate, 

 about as long (20 mm.) as the spurs. The sepals are mainly 



