12 



with the somites. The mandibles are weil developed; the inner lobes of 

 the maxillipeds are fused. 



Depending on the structure of the antennae, this tribe was divided 

 by Woltereck into three subtribes: Hyperiidea Recticornia, Hyperiidea 

 Filicornia, and Hyperiidea Curvicornia. Schellenberg (1942) included in 

 the group Genuina only the first two subtribes; the third he indentified 

 as a special tribe, Hyperiidea Anomala. 



Pirlot (1929) justifiably noted that the hierarchy proposed by 

 Woltereck is rather artificial and not at all acceptable. Actually, the 

 genera Mimonecteola and Microphasma from the group Primitiva, which 

 Woltereck considered the most primitive, are distinctly neotenic, and hence 

 cannot stand at the beginning of the evolutionary series of Hyperiidea. 

 The phyletically very close families Lanceolidae and Chuneolidae fall 

 into different groups in this system — Completa and Incompleta; the 

 family Mimonectidae has been included in the subtribe Primitiva, and 

 the rather closely related family Scinidae in the subtribe Derivata. The 

 same subtribe includes the family vibiliidae, sharply differing from the 

 family Scinidae and from other Gammaridea. On the other hand, the 

 family Cyllopodidae, which is extremely close to the Vibiliidae, has been 

 included in the tribe Genuina. Finally, in recent times, in the family 

 Hyperiidae (tribe Genuina) the genus Bougisia has been included, with 

 a small head and very small eyes, which would necessitate changing the 

 diagnosis of the tribe. Pirlot proposed a more natural classification of the 

 suborder Hyperiidea. He divided the suborder into two tribes: Hyperiidea 

 Physosomata and Hyperiidea Eugenuina (Physocephalata according to 

 Bowman and Gruner, 1973). In the former tribe, whose characteristics 

 21 are given below, two groups are distinctly differentiated. The first is 

 characterized by mandibles with a broad cutting edge, comparatively short 

 antennae I, strong pereopods I with distally enlarged 5th segment, and 

 comparatively large coxal plates; it combines the families Microphasmidae, 

 Lanceolidae, and Chuneolidae. The genus Mimonecteola of the family 

 Microphasmidae is the least specialized; other genera of this family are 

 more obligate commensals or parasites. Intensification of the obligate 

 nature of parasitism of coelenterates proceeded along the line Lanceolidae- 

 Chuneolidae. The latter have obviously more completely lost the ability 

 to swim freely and lost the mandibular palp but, contrarily, have acquired 

 retractile claws on all the pereopods, ensuring a more complete attachment 

 to the host tissue. Stephensen and Pirlot ( 193 1 ) combined all these families 

 in the group Lanceoliformata. 



The second group— Sciniformata characterized by a long, conical, 

 fairly thin proximal article of the flagellum of antennae I, distinct sex- 

 ual dimorphism in the structure of antennae II, mandibles with a narrow 

 cutting edge and without a mandibular palp, and almost not enlarged, 



