Claw of anal legs strong and strongly curved. 



Pupa. Length of male pupae 11 mm, of female pupae 13 mm. Antennae 

 filiform, thin, reaching middle of segment 6 in male pupae, segment 5 in 

 female pupae; 1st segment larger than the others; labrum broad, with 

 slightly rounded anterior margin and indistinctly marked angles; each 

 angle with a long seta; 3 such setae and a longer seta laterally at the 

 anterior margin. Mandibles strong and long, curved at the base, with 

 3 apical teeth; teeth and blades finely serrated; outer setae almost of the 

 same length. Maxillary palps 5 -segmented; terminal segment longer than 

 all the other segments together; labial palps 3 -segmented; the last segment 

 as long as the two preceding segments together. Wing sheaths short and 

 pointed, of different length, second pair reaching the middle of abdominal 

 segment 3. Mid-tarsi with blackish brown, dense natatorial setae and strong 

 claws. Holding apparatus of segments 3—7 weakly developed; number of 

 spinules as follows: 111:3—6; IV:3— 7; V:3— 5+ numerous minute spinules on 

 the postsegmental plates; VI: 6— 9. The long ventral appendages of the male 

 genitalia are divided to the middle; penis sheath short. 



Mode of life and habitats. Brooks and rivulets with cool water 

 and rapid current. The record of Ph. montanus from the Molozhskii 

 area of the Rybinsk Reservoir is doubtful. 



Distribution. Northwest and west of the European USSR. In 

 addition, Europe. 



3. Genus Dolophilodes Ulmer 



Full-grown larva. Length about 14—15 mm. Head oblong, almost twice 

 as long as pronotum in some Middle Asian species, brownish or chestnut 

 brown, without pattern; frontoclypeus asymmetrical anteriorly; gula short, 

 broadly triangular, partly projecting beyond the head capsule. Eyes small, 

 situated close to the anterior margin. Labrum with a weak, light dorsal 

 triangular sclerite in the middle; anterior membranous margin of labrum 

 straight; row of minute hairs on it continuous; median indentation hardly 

 marked. Mandibles large and strong, black or blackish brown, with an 

 undulate outer surface, and with two light setae, which are widely separated 

 and situated at a marked distance from the basal margin. Pronotum and 

 head brownish or chestnut brown, with straight lateral margin; posterior 

 angles slightly prolonged; a broad and continuous black border at the 

 posterior and lateral margin; margin light and milk white posteriorly; 

 posterior area of sclerite broader in the middle than laterally. Trochantin 

 prolonged in a horn, shorter than in Wormaldia and Philopotamus. 



Legs of almost the same length; forelegs slightly darker than the other 

 legs; hind legs shorter and thinner. Anterodorsal seta of fore coxa dark, 

 spinelike, resembling the basal seta of the coxa. Tarsi of forelegs with a 

 continuous row of small light ventral spinules. Similar spinules in the 

 distal part of mid- and hind tarsi. Claws with a process near the long 

 light seta. Abdominal segments (except segment 9) uniformly broad, as in 

 the other genera; claws of anal legs dark. This genus has been little studied.* 



* In addition to the larva described below (from the vicinity of Lake Teletskoe), the collections of the 

 Zoological Institute also contain several larvae of Dolophilodes of an unidentified species from 

 Kirghizia and Tadzhikistan; these specimens were used for the description of the genus. 



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