DEVELOPMENT OF SPERMATOBIUM. 21 



Frequently we find a restkorper consisting of a num- 

 ber of unused cytospheres scattered about, but princi- 

 pally situated in the center of the sporoblast. I think 

 that the name sporophore is an unsuitable one for this 

 body, because, as has frequently been remarked, the 

 restkorper consists simply of the unused cytospheres, 

 which perhaps for want of nuclei have not consolidated 

 into sporoblasts. 



THE SHUTTLE SPORES AND PSEUDONAVICELL^. 



The ultimate transformation of the sporoblasts are into 

 shuttle spores and later the pseudonavicell«. At the 

 •earliest stage these bodies are, however, lunate, crescent- 

 shaped, concave, considerably varying in form, but not 

 in size. Each one contains a yellow staining crescent, 

 possibly the accumulation of nuclear matter, always sit- 

 uated close to the convex side (figs. 24 and 25). 



I have never been able to clearly make out the struc- 

 ture of the spores at this stage, and, while assuming that 

 they really are only earlier stages of the shuttle spores, 

 I have yet some hesitation as to this being the case ; the 

 reason for this is that I have never distinctly seen in them 

 the cytospheres, which, however, are always very distinct 

 in the shuttle and pseudonavicellas spores. 



One-half or more of the lunate spore is occupied by a 

 transparent lunate sac, while the other half or less con- 

 cists of a granulated crescent. 



The real shuttle spores are more regular, but even they 

 show a concave side, but their contents can always be 

 clearly dissolved into cytospheres and micronuclear bodies 

 of varying number (figs. 32, 33, etc.) This is their 

 shape while yet enclosed in the sporogonium. Their 

 cytothecas are there thin, hardly visible, but always well 

 defined. Such shuttle spores are also found free among 

 the spermatozoa of the host, and show these often irre- 



