Nr. 11 ANATOMICAL STUDIES ON ANELASMA AND SCALPELLUM 19 



(1904, 1905). They laek brown granulations, and should accoird- 

 ingly rather approach the pancreatic group, but the minute 

 structure of the cells differs generally from the pancreatic oells as 

 described by Gruvel. Their function is probably a mixed one, 

 and although they do not completely correspond with Gruvél's 

 descriptions, they must be ranged aimong the hepatico-panoreatic 

 group. We are at present incapable of discerning the chemical 

 character of the glands in question, the only safe basis for a 

 elassification in the categories named ahove. 



Finally, some details as to the c e m e n t a r y glan d s of 

 Scalpellum Stromii may be given here. The oementary glands are 

 situated at the upper (hinder) side of the ovaries, under the «stalk» 

 or connecting part between the thorax and the peduncle. The 

 glands consiist of a series of large gland cells connected by fine 

 ducts like the grapes of a cluster (pl. III, fig. 16); the fine ducts 

 unite into one main duct leading down the peduncle, and opening 

 out at the base of the same. The animal has two such composite 

 glands, one on each side of the sagittal plane; there are iaccordingly 

 al so two symmetrically placed main ducts in the peduncle. — The 

 single gland cell has a finely granulated protoplasm; the granula 

 exhibit a rather strong affinity to Delafields haematoxyline, and 

 might thus be judged as mucous; staining with mucicarrnine 

 nevertheless completely fails. Whereas the granula as mentioned, 

 are basophile, the cytoplasm is otherwise pronouncedly eosinophile. 

 The nuclei are extraordinarily large, and irregularly amoeboid 

 shaped, generally lobed. The chromatic substance appaars as 

 numerous, densely crowded small grams, which are characterized 

 by their pronounced affinity to haeniatoxy lines and boraxcarmine; 

 Delap^ield's haematoxyline stains them blackish blue. The latter 

 dye also brings to light one or two large nucleoli in the nucleus; 

 also in the oementary gland oells the nucleoli are at the same time 

 eosinophile. The nucleoli are also distinct in preparations stained 

 with boraxcarmine. 



The cementary gland oells are evidently in vivid function; 

 nevertheless their secretions could not be made out clearly in the 

 ducts. There must be a difference in the nature of the function 

 of the cementary glands in Scalpellum and Anelasma; this is 

 clearly shown by their different relation to a double staining with 

 boraxcarmiine — bleu de Lyon. In Scalpellum, the cytoplasm of the 

 cementary gland cells attains a blueish violet hue, whereas the 

 cytoplasm of the same cells in Anelasma (pl. V, fig, 24) attains an 

 almost pure red colour and, acoorddngly, turns out to be more 

 strongly basophile than in Scalpellum. 



