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•DNTTED STATES NATIONAL MUSEUM BULLETIN 215 



wMch are smooth and centrally excavated, attached in 

 the later stages, with the chambers slightly inflated, 

 extremely papUlose and fistulose although retaining 

 somewhat the chevron-shaped character; sutures raised 

 and thickened in the early portion, slightly depressed 

 in the irregidar attached portion; wall calcareous, 

 perforate; aperture terminal like Citharinella in the 

 early stages, with numerous apertures at the ends of 

 the fistulose extensions in the later attached chambers. 

 Remarks: Tentifrons, new genus, diEfers from all 

 palmate Nodosariidae in being attached in its later 

 stages, in developing the fistulose growth, and in having 

 multiple apertures. These characters show again the 

 close aflBnity between the families Nodosariidae and 

 Polymorphinidae. 



Tentifrons bamardi Loeblicb and Tappan, new species 



Plate 72, Figures 17, 18 



FldbelUna cf. angulosa d'Orbigny Barnard, 1949 (not Frondicu- 

 laria angulosa d'Orbigny, 1840), Proc. Geol. Assoc, vol. 60, 

 pt. 4, p. 285, pi. 12, figs. 1-6. 



Test large, flattened, free in the early stage, later 

 attached; chambers increase gradually in size from the 

 globular proloculus, at first in a citharine arrangement, 

 later uniserial with chevron-shaped chambers, and 

 flattened or slightly depressed, the final chambers 

 formed after the test becomes attached are irregular in 

 outline, more inflated, and with a distinctly papillose 

 surface and some develop numerous fistulose extensions 

 along the chamber margins; sutures raised and thick- 

 ened in the early stages, formed during the free develop- 

 ment, but slightly depressed between the later fistulose 

 attached chambers; wall calcareous, perforate, surface 

 smooth in the early free portion, distinctly ornamented 

 in the later portion; aperture terminal, as in Citharinella, 

 at first at the dorsal angle, then terminal and central, 

 but in the later attached portion there are numerous 

 apertures at the ends of the fistulose extensions of the 

 chambers along their margins. 



Length of early free stage of holotype 1.4 mm., 

 greatest breadth 0.84 mm., total length of test 8.9 mm., 

 greatest breadth of fistulose chambers 2.99 mm. 



Remarks: Barnard (1949, p. 285) described these 

 specimens under the name Flabellina cf. angulosa 

 d'Orbigny. Evidently this combination of names must 

 be in error as d'Orbigny described no species F. 

 angulosa so far as the authors have been able to deter- 

 mine. In addition no species of Cristellaria was 

 termed angulosa by d'Orbigny. The species Frondicu- 

 laria angulosa d'Orbigny is so different that it could not 

 be this species. As no parentheses were placed around 

 d'Orbigny's name, the writers are in doubt as to what 

 species Barnard referred these peculiar forms from the 

 English Chalk. However, the early stage of the present 

 species compares favorably with that of Cristellaria 

 gaudryana d'Orbigny, 1840. 



Barnard considered these forms to be abnormal 

 specimens, evidently "freaks"; but they seem to repre- 



sent a trend in the development of the palmate Nodo- 

 sariidae, the end stage of one line of evolution. Cer- 

 tainly such forms, evidently not rare, which change 

 from a free mode of existence to a fixed life, and develop 

 a fistulose end stage with numerous apertures are 

 worthy of recognition as a distinct genus. 



Our illustration of the holotype (pi. 73, fig. 18), 

 shows two more chambers than the illustration of this 

 specimen published by Barnard (1949, pi. 12, fig. 6); 

 however, these last two chambers are somewhat 

 abraded, and are represented largely by a mere outline 

 and some fragmentary portions of the test along the 

 margins. 



Tentifrons bamardi, new species, is similar to Cristel- 

 laria gaudryana d'Orbigny in size and shape of the 

 early test, but C. gaudryana lacks any tendency to 

 develop the fistulose growth in the later stage, and does 

 not show the pustulose wall and inflated later chambers 

 also typical of the present species. 



Types and occurrence : Holotype (BMNH P40275), 

 specimen figured by Barnard (1949), pi. 12, fig. 6 and 

 paratype (BMNH P40274), specimen figured by 

 Barnard (1949), pi. 12, fig. 5, both from the Belem- 

 nitella mucronata zone (Upper Senonian) of the Upper 

 Chalk of Tharston, Norfolk County, England. Col- 

 lected by A. W. Rowe. 



Barnard (1949, pi. 12) also figured specimens similar 

 to these from Councils Pit, Newmarket Road, Norwich, 

 and from StonehUl Kiln, Norwich, both also from the 

 Belemnitella mucronata zone of the Upper Chalk 

 (BMNH P40272 and P40273). 



Family Polymorphinidae d'Orbigny, 1846 

 Genus Webbinella Rhumbler, 1904, emended 



Webbinella RHtFMBLBE, Archiv. Protistent., Band 3, Teil 1, p. 

 228, 1904. 



Plate 72, Figure 19 



Type species: WebbinM hemisphaerica Jones, Parker 

 and Brady, 1865. Subsequent designation by Cushman 

 (1918, p. 61). 



Test attached, consisting of an initial polymorphine 

 stage of three chambers, surrounded by a larger circular 

 fourth chamber spreading on the surface of the sub- 

 stratum; wall calcareous, perforate; aperture not 

 evident. 



Remarks: The holotype of Jones, Parker and Brady, 

 in the British Museum, is an attached hemispherical 

 form, but is calcareous and not agglutinated. Further- 

 more, when dampened with glycerine the chamber 

 divisions can be seen and the species is shown to be a 

 polymorphinid, completely unlike the arenaceous forms 

 usually placed under Webbinella. It is similar therefore 

 to Bullopora Quenstedt, but differs in having an early 

 multUocular polymorphine stage. It differs from 

 Histopomphus Loeblich and Tappan in having a circuJar 

 spreading attachment and not a branched and 

 irregular one. 



