AMERICAN MOTHS OF THE SUBFAMILY PHTCITINAE 



99 



53. Genus Etiella Zeller 



Eiiella Zeller, Isis von Oken, 1839, p. 179; 1846, p. 733.— Heine- 

 mann, Die Schmetterlinge Deutschlands und der Schweiz, 

 Abt. 2, vol. 1, pt. 2, p. 154, 1865.— Meyrick, Proc. Linn. 

 Soc. New South Wales, vol. 3, p. 629, 1882.— Hulst, Phy- 

 citidae of N. Amer., p. 169, 1890; U. S. Nat. Mus. Bull. 52, 

 p. 428, 1902.- Ragonot, Monograph, pt. 1, p. 569, 1893.— 

 Spuler, Die Schmetterlinge Europas, vol. 2, p. 208, 1910. — 

 Forbes, Cornell Mem. 68, p. 629, 1923.— Janse, Journ. Ent. 

 Soc. South Africa, vol. 7, p. 15, 1944. (Type of genus: 

 Phycis zinckenella Treitschke.) 



Ramphodes Gu6n6e, Europaeorum Microlepidopterorum index 

 methodicus . . ., p. 81, 1845. (Type of genus: Phycis 

 zinckenella Treitschke.) 



Tongue well developed. Antenna of male with basal 

 segment enlarged and bearing a short, bluntly pointed 

 projection on inner side near base; shaft with sinus and 

 large hair and scale tuft at base, pubescent; antenna of 

 female simple. Labial palpus porrect; very long (the 

 length of head and thorax); smoothly scaled; second 

 segment about five times the length of third on male 

 and grooved to hold maxillary palpus; third segment 

 short on male, longer on female, acuminate, frequently 

 bent downward, especially on female. Maxillary palpus 

 of male in the form of an aigrette; of female small, 

 squamous. Forewing with ridge of raised scales beyond 

 base; 11 veins; vein 2 from before but near lower outer 

 angle of cell; 3 from the angle, approximately equi- 

 distant from 2 and 4 ; 4 and 5 shortly separated at base, 

 diverging from cell; 6 from below upper angle of cell, 

 straight; 8 and 9 stalked for half their lengths; 10 from 

 the ceU, separated and divergent from the stalk of 8-9 ; 

 male without costal fold. Hind wing with vein 2 from 

 well before lower outer angle of cell; 3 from the angle, 

 appreciably shorter than 2, closely approximate to the 

 stalk of 4-5 at base (separated by a very short vein) ; 

 4 and 5 normally stalked (rarely weakly anastomosed 

 or contiguous) to about middle; 7 and 8 closely approxi- 

 mate for less than haK their lengths from cell; cell about 

 one-third the length of wing on male, slightly longer on 

 female; discocellular vein curved. Eighth abdominal 

 segment of male with a pair of weak ventrolateral hair 

 tufts. 



Male genitalia with uncus hoodlike, its apical margin 

 evenly rounded. Apical process of gnathos a simple, 

 sharp hook. Transtilla incomplete and vestigial (its 

 elements rarely distinguishable except under high 

 magnification). Harpe with a strongly sclerotized, 

 curved, tapering, pointed arm projecting from base of 

 costa and as long as costa; remainder of harpe wealdy 

 sclerotized and abruptly narrowed at middle (the shape 

 of harpe probably a specific character). Anellus V- 

 shaped; its arms rather broad, blunt and haired. 

 Aedeagus moderately long, stout; penis armed with 

 two strong cornuti. Vinculum stout ; as long as greatest 

 width; evenly tapering to a blunt point. 



Female genitaha with bujsa elongate, finely scobinate, 

 armed with signa consisting of curved, sclerotized bands 

 armed with slender spines and situated near junction of 

 bursa and ductus bursae ; a sclerotized lobe on bursa near 

 ductus bursae; ductus bursae short (much shorter than 



bursa), tubular, sclerotized, ribbed and broadened 

 towards genital opening; ductus seminalis from bursa 

 adjacent to sclerotized lobe. 



A distinct genus, not to be confused with anything 

 else and easily identified by its combination of antennal, 

 palpal, wing, and genitalic characters. Ragonot, Hulst, 

 and Janse list five Walker names in the generic synony- 

 my. These supposed genera were based on Old World, 

 tropical species which have all been referred as s3tio- 

 njins of zinckenella. I have no reason to question this 

 synonymy but have omitted the references as I have 

 not been able to check the genitalia or their types. The 

 only species occiu-ring in the New World is zinckenella. 



The larva differs from typical phycitid larvae in that 

 it lacks altogether the sclerotized rings about seta lib 

 of mesothorax and seta III of the eighth abdominal 

 segment. 



207. Etiella zinckenella (Treitschke) 

 Figures 17, 326, 840 



Phycis zinckenella Treitschke, Die Schmetterlinge von Europa, 

 vol. 9, pt. 1, p. 201, 1832. 



Phycis etiella Treitschke, Die Schmetterlinge von Europa, vol. 

 10, p. 3, p. 174, 1835. — Duponchel, Histoire naturelle des 

 l^pidoptSres, ou papillons de France, vol. 10, p. 180, 1836. — 

 Milliere, Iconographie et description de chenilles et l§pidop- 

 tfires inedits, vol. 1, p. 248, 1861. (Originally proposed as 

 new name for zinckenella.) 



Pempelia Eiiella zinckenella (Treitschke) Zeller, Isis von Oken, 

 1839, p. 179; 1846, p. 755.— Herrioh-Schafifer, Systematische 

 Bearbeitung der Schmetterlinge von Europa, vol. 4, p.^'72, 

 1849. 



Ramphodes zinckenella (Treitschke) Gu^n^e, Europaeorum 

 Microlepidopterorum index methodicus . . . , p. 81, 1845. 



Eiiella zinckenella (Treitschke) Heinemann, Die Schmetterlinge 

 Deutschlands und der Schweiz, Abt. 2, vol. 1, pt. 2, p. 154, 

 1865. — Zeller, Horae Soc. Ent. Rossicae, vol. 16, p. 177, 

 1881.— Hulst, Phycitidae of N. Amer., p. 170, 1890; U. S. 

 Nat. Mus. Bull. 52, p. 428, 1902.— Ragonot, Monograph, 

 pt. 1, p. 572, 1893.— Chittenden, U. S. Dep. Agr. Bur. Ent. 

 Bull. 82 (pt. 3), p. 25, 1909. — Essig, Insects of western 

 North America, p. 709, 1926. — Walcott, Journ. Agr. Univ. 

 Puerto Rico, vol. 20, no. 1, p. 476, 1936.— McDunnough, 

 Check list. No. 6274, 1939. 



Etiella zinckenella schisticolor Zeller, Horae Soc. Ent. Rossicae, 

 vol. 16, p. 178, 1881.— Hyslop, U. S. Dep. Agr. Bur. Ent. 

 Bull. 95, pt. 6, p. 82, 1912. 



Etiella villosella Hulst, Ent. Amer., vol. 3, p. 133, 1887. 



Etiella schisticolor (Zeller) Ragonot, Ent. Amer., vol. 5, p. 116, 

 1889; Monograph, pt. 1, p. 274, 1893.— Hulst, Phycitidae 

 of N. Amer., p. 170, 1890. 



Etiella rubribasella Hulst, Phycitidae of N. Amer., p. 170, 1890. — 

 Ragonot, Monograph, pt. 1, p. 572, 1893. 



Forewing gray more or less lightened by white 

 scaling, especially in median areas; a broad white band 

 along costa, extending from base to or nearly to apex; 

 extreme costal edge dark gray to red; on occasional 

 specimens some red scaling at extreme base of wing and 

 in outer median area; transverse lines obsolete; the 

 antemedial line replaced by a ridge of raised scales near 

 basal third and extending from inner margin to top of 

 cell, the raised scales metallic ocherous or orange red 

 bordered outwardly by a broad ocherous or orange 

 patch; discal spots obsolete. Hind wing whitish with 

 a faint smoky tint, to dark smoky fuscous; the veins 



