228 



tmiTED STATES NATIONAL MUSEUM BUTiLETIN' 207 



well separated from 8-9; male with a short costal fold. 

 Hind wing with vein 2 from well before angle of cell; 3 

 from before but near angle, approximate to 5; 6 from 

 near upper angle of cell; 7 and 8 completely fused be- 

 yond cell (in occasional specimens a rudiment of 8 visible 

 as a short spur from outer fifth of vein 7) ; discocellular 

 vein straight and vertical; cell approximately one-third 

 the length of the wing. Eighth abdominal segment of 

 male simple. 



Male genitalia with uncus bx'oad, subtriangular, 

 outer surface densely covered with bristlelike scales. 

 Gnathos terminating in a moderately large, fused or 

 partially fused, hooked apical process. Harpe simple; 

 costa strongly sclerotized for most of its length; apex 

 broadly rounded. Anellus U-shaped; lateral arms 

 curved and partially encircling aedeagus. Aedeagus 

 straight; somewhat constricted at apex; smooth or 

 with a patch of minute scobinations at apex. Penis 

 with some scobinate wrinklings, otherwise imarmed. 

 Vinculimi stout and broad, long and tapering slightly 

 to more or less narrowly roimded terminal margin. 



Female genitalia with bursa copulatrix membranous, 

 finely scobinate over caudal half of inner surface; 

 signum consisting of a girdle of fine, narrow, serrate, 

 ridgelike disks about middle of bursa, or completely 

 absent. Ductus bursae membranous throughout, finely 

 scobinate towards junction with bursa. Genital open- 

 ing unsclerotized. Ductus seminalis from bursa near 

 its junction with ductus bursae. 



Richards and Thomson (1932) referred Strymax 

 Dyar to Ephestia as a subgenus. They did not treat 

 Unadilla. Hulst erected the latter for a single species 

 {nasutella Hulst) which Hampson placed with two other 

 Zeller species (ubacensis and maturella) in the synonymy 

 of erronella Zeller. Upon the basis of this synonymy, 

 which is doubtful, to say the least, Hampson cites 

 erronella as the type of Unadilla. Janse follows him 

 in this. Nomenclatorily nasutella must be the type of 

 Unadilla whether nasutella proves to be a synonym of 

 erronella or not. 



Strymax is an obvious synonym of Unadilla. It has 

 no close affinity to Ephestia even in the broad sense in 

 which that genus is interpreted by Richards and 

 Thomson. Hind wing venation and genitalia seem to 

 indicate much closer relation to Homoeosoma. 



On structures of the male and female genitalia the 

 species divide into two distract groups, as follows: 



Male with apical process of gnathos partially fused; female 



with an encircling band of signa. 

 Male with apical process of gnathos completely fused; 



female without signa. 



Genus Unadilla, Species 469: U. erronella 



[Male with apical process of gnathos partially fused; female 

 with an encircling band of signa.] 



469. Unadilla erronella (Zeller) 



Figures 121, 498, 995 



Homoeosoma erronella Zeller, Horae Soc. Ent. Rossicae, vol. 16, 

 p. 238, 1881. 



Homoeosoma ubacensis Zeller, Horae Soc. Ent. Rossicae, vol. 16, 



p. 239, 1881. 

 Unadilla erronella (Zeller) Hampson, in Ragonot, Monograph, 



pt. 2, p. 262, 1901. 

 Ephestia bipunciella Hampson, Ann. Mag. Nat. Hist., ser. 7, vol. 



7, p. 255, 1901 (new synonymy). 

 Strymax dorae Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 344, 1914 



(new synonymy). 

 Strymax pyllis Dyar, Proc. U. S. Nat. Mus., vol. 47, p. 344, 1914 



(new synonymy). 

 Ephestia (Strymax) bipunctella (Hampson) Richards and Thom- 

 son, Trans. Ent. Soc. London, vol. 80, p. 201, 1932. 

 Ephestia {Strymax) pyllis (Dyar) Richards and Thomson, 



Trans. Ent. Soc. London, vol. 80, p. 202, 1932. 



Hampson referred ubacensis as a synonym of erronella, 

 and, I believe, correctly so, judging from photographs 

 of the male types and their genitalia. The only differ- 

 ence between the two is in the more intensified macu- 

 lation of ubacensis. The same difference is exhibited 

 by the male types of dorae and pyllis, the latter being a 

 pale example with the normal subbasal markings of 

 forewing almost obsolete. However, in a series of 

 dorae from Panamd there are several intergrading 

 examples. U. erronella is also variable in size and to 

 some extent in structure, the vinculum being shorter in 

 some specimens from Puerto Rico than in those from 

 Panamd and Colombia, but otherwise the genitalia are 

 remarkably constant. In careful preparations of the 

 genitalia of Dyar's supposed two species (dorae and 

 pyllis) I am unable to find any of the differences shown 

 in the figures pubUshed by Richards and Thomson. 

 The general color of the forewing is whitish gray; sub- 

 basal markings, when present, consisting of from one to 

 three pale brownish spots forming a broken antemedial 

 transverse shade; discal dots faint; hind wing white in 

 the male, more or less shaded with pale smoky fuscous 

 in the female. Alar expanse, 9-14 mm. 



Type localities: Honda, Colombia (erronella, in 

 BM); Ubaque, Colombia (ubacensis, in BM); Nassau, 

 Bahamas (bipunctella, in BM); La Chorrera, Panami 

 (dorae, in USNM); Porto BeUo, Panamd (pyllis, in 

 USNM). 



Food plant: Unknown. 



Disteibution: Colombia; Honda, Mariquita, 

 Ubaque. PanamX: Corazal (Mar., Apr., May), La 

 Chorrera (Apr., May), Paraiso (Apr.), Porto Bello 

 (Feb.), Rio Trinidad (Mar., May), Taboga Isl. (Feb., 

 June). Bahamas: Nassau. Puerto Rico: Aguu-re 

 Central (Apr.), Catano (Apr., June), Coamo Springs 

 (Apr.), Dorado (May, Jime), Isabela (Apr.), San 

 German (Aug.); Puerto Real (Vieques Isl., Apr., July). 

 Virgin Islands: Kingshill (St. Croix, Mar., Dec). 



The Puerto Rican specimens (some 40-odd in the 

 Cornell University Collection) may possibly represent 

 a distinct race from the mainland form, but I am 

 imable to find any valid character for their separation. 

 The somewhat shorter vinculum of the male is matched 

 in some Panamd specimens and is of very doubtful 

 significance. U. erronella is easily identified as it is 

 the only described species with a partially divided 

 apical process of gnathos and a girdle of signa. The 



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